Professor Sean Connell

Sean Connell
School of Biological Sciences
Faculty of Sciences

Sea creatures, their life and their future - is my work. This daily life has inspired new policy & cleaner coasts by scientific discovery & saner theory.

Join me. I'm looking for students interested in biology and marine biology (PhD, Honours & MSc students). All projects are possible. Those underway include climate change and restoration ecology (e.g. restoring 20 hectares of oyster reef). Students snorkel & dive or use mesocosms & aquariums. These techniques enable a variety of projects - from behaviour through tests of how individuals work together to form communities.

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External Profiles

Professor Sean Connell

Sea creatures, their life and their future - is my work. This daily life has inspired new policy & cleaner coasts by scientific discovery & saner theory.

Join me. I'm looking for students interested in biology and marine biology (PhD, Honours & MSc students). All projects are possible. Those underway include climate change and restoration ecology (e.g. restoring 20 hectares of oyster reef). Students snorkel & dive or use mesocosms & aquariums. These techniques enable a variety of projects - from behaviour through tests of how individuals work together to form communities.

Add these ingredients (below); work, people, fun, adventure and data...

Sean @ Work
       Sean @ Work
  With strangers
           Having fun
        In exotic locations
Sean writing down hard won data
    Getting data

... to get results (below)...

Collecting data in exotic locations with strangers who are fun to work with.  For students, visit my personal website and supervision tab on this page.

a a Book
a a a







Date Position Institution name
2011 Professor University of Adelaide
2005 - 2010 Associate Professor University of Adelaide
1999 - 2004 Senior Lecturer The University of Adelaide
1997 - 1999 Lecturer University of Sydney
1996 - 1997 Lecturer University of Tasmania (Australian Maritime College)
1994 - 1996 Scientist The Ecology Laboratory

Awards and Achievements

Date Type Title Institution Name Amount
2010 Fellowship ARC Future Fellowship
2004 Fellowship ARC Queen Elizabeath II Fellowship
2003 Award J.G. Russell Award Australian Academy of Science
2003 Award The Andrewartha Medal
1992 Research Award Scholarship (PhD) - Australia University of Sydney
1990 Award Scholarship (PhD) - New Zealand New Zealand University Grants Committee


Date Institution name Country Title
University of Auckland New Zealand MSc (1st Class Honours)
University of Sydney Australia PhD
University of Canterbury New Zealand BSc

Research Interests

Aquatic Ecosystem Studies and Stock Assessment, Climate Change Processes, Community Ecology, Ecological Impacts of Climate Change, Ecology, Ecosystem Function, Marine and Estuarine Ecology


Year Citation
2017 Connell, S., Fernandes, M., Burnell, O., Doubleday, Z., Griffin, K., Irving, A. ... Falkenberg, L. (2017). Testing for thresholds of ecosystem collapse in seagrass meadows. Conservation Biology, 31, 5, 1-6.
2017 Rossi, T., Connell, S. & Nagelkerken, I. (2017). The sounds of silence: regime shifts impoverish marine soundscapes. Landscape Ecology, 32, 2, 239-248.
2017 Hardy, N., Berry, T., Kelaher, B., Goldsworthy, S., Bunce, M., Coleman, M. ... Figueira, W. (2017). Assessing the trophic ecology of top predators across a recolonisation frontier using DNA metabarcoding of diets. Marine Ecology Progress Series, 573, 237-254.
2017 Sunday, J., Fabricius, K., Kroeker, K., Anderson, K., Brown, N., Barry, J. ... Harley, C. (2017). Ocean acidification can mediate biodiversity shifts by changing biogenic habitat. Nature Climate Change, 7, 1, 81-85.
2017 Pistevos, J., Nagelkerken, I., Rossi, T. & Connell, S. (2017). Antagonistic effects of ocean acidification and warming on hunting sharks. Oikos, 126, 2, 241-247.
2017 Vizzini, S., Martinez-Crego, B., Andolina, C., Massa-Gallucci, A., Connell, S. & Gambi, M. (2017). Ocean acidification as a driver of community simplification via the collapse of higher-order and rise of lower-order consumers. Scientific Reports, 7, 1, 4018-1-4018-10.
2017 Harley, C., Connell, S., Doubleday, Z., Kelaher, B., Russell, B., Sarà, G. & Helmuth, B. (2017). Conceptualizing ecosystem tipping points within a physiological framework. Ecology and Evolution, 7, 15, 1-11.
2017 Leung, J. Y., Connell, S. D., Nagelkerken, I. & Russell, B. D. (2017). Impacts of Near-Future Ocean Acidification and Warming on the Shell Mechanical and Geochemical Properties of Gastropods from Intertidal to Subtidal Zones.. Environmental Science & Technology, -.
2017 Doubleday, Z. & Connell, S. (2017). Publishing with Objective Charisma: Breaking Science’s Paradox. Trends in Ecology and Evolution, -.
2017 Pistevos, J., Nagelkerken, I., Rossi, T. & Connell, S. (2017). Ocean acidification alters temperature and salinity preferences in larval fish. Oecologia, 183, 2, 545-553.
2017 Provost, E., Kelaher, B., Dworjanyn, S., Russell, B., Connell, S., Ghedini, G. ... Coleman, M. (2017). Climate-driven disparities among ecological interactions threaten kelp forest persistence. Global Change Biology, 23, 1, 353-361.
2017 Goldenberg, S., Nagelkerken, I., Ferreira, C., Ullah, H. & Connell, S. (2017). Boosted food web productivity through ocean acidification collapses under warming. Global Change Biology, 23, 10, 4177-4184.
2017 Ghedini, G. & Connell, S. (2017). Moving ocean acidification research beyond a simple science: Investigating ecological change and their stabilizers. Food Webs, Online, 1-7.
2017 Connell, S., Doubleday, Z., Hamlyn, S., Foster, N., Harley, C., Helmuth, B. ... Russell, B. (2017). How ocean acidification can benefit calcifiers. Current Biology, 27, 3, R95-R96.
2017 Nagelkerken, I., Goldenberg, S., Ferreira, C., Russell, B. & Connell, S. (2017). Species interactions drive fish biodiversity loss in a high-CO₂ world. Current Biology, 27, 14, 2177-2184.e4.
2017 Leung, J., Russell, B. & Connell, S. (2017). Mineralogical plasticity acts as a compensatory mechanism to the impacts of ocean acidification. Environmental Science and Technology, 51, 5, 2652-2659.
2017 Davis, K. L., Coleman, M. A., Connell, S. D., Russell, B. D., Gillanders, B. M. & Kelaher, B. P. (2017). Ecological performance of construction materials subject to ocean climate change.. Marine environmental research, -.
2016 Heldt, K., Connell, S., Anderson, K., Russell, B. & Munguia, P. (2016). Future climate stimulates population out-breaks by relaxing constraints on reproduction. Scientific Reports, 6, 1, 33383-1-33383-8.
2016 Rossi, T., Nagelkerken, I., Pistevos, J. & Connell, S. (2016). Lost at sea: ocean acidification undermines larval fish orientation via altered hearing and marine soundscape modification. Biology Letters, 12, 1, 1-4.
2016 Falkenberg, L., Russell, B. & Connell, S. (2016). Design and performance evaluation of a mesocosm facility and techniques to simulate ocean acidification and warming. Limnology and Oceanography: Methods, 14, 4, 278-291.
2016 Alleway, H., Gillanders, B. & Connell, S. (2016). 'Neo-Europe' and its ecological consequences: the example of systematic degradation in Australia's inland fisheries. BIOLOGY LETTERS, 12, 1, 1-7.
2016 McSkimming, C., Connell, S. D., Russell, B. D. & Tanner, J. E. (2016). Habitat restoration: early signs and extent of faunal recovery relative to seagrass recovery. Estuarine, Coastal and Shelf Science, 171, 51-57.
2016 Connell, S., Nimmo, D., Ghedini, G., Mac Nally, R. & Bennett, A. (2016). Ecological Resistance - Why Mechanisms Matter: A Reply to Sundstrom et al.. Trends in Ecology and Evolution, 31, 6, 413-414.
2016 Ghedini, G. & Connell, S. (2016). Organismal homeostasis buffers the effects of abiotic change on community dynamics. Ecology, 97, 10, 2671-2679.
2016 Rossi, T., Connell, S. & Nagelkerken, I. (2016). Silent oceans: ocean acidification impoverishes natural soundscapes by altering sound production of the world’s noisiest marine invertebrate. Proceedings of the Royal Society B, 283, 1826, 20153046-1-20153046-7.
2016 Krumhansl, K., Okamoto, D., Rassweiler, A., Novak, M., Bolton, J., Cavanaugh, K. ... Edgar, G. (2016). Global patterns of kelp forest change over the past half-century. Proceedings of the National Academy of Sciences of the United States of America, 113, 48, 13785-13790.
2016 Nagelkerken, I., Russell, B., Gillanders, B. & Connell, S. (2016). Ocean acidification alters fish populations indirectly through habitat modification. Nature Climate Change, 6, 1, 89-93.
2015 Rossi, T., Nagelkerken, I., Simpson, S., Pistevos, J., Watson, S., Merillet, L. ... Connell, S. (2015). Ocean acidification boosts larval fish development but reduces the window of opportunity for successful settlement. Proceedings of the Royal Society B, 282, 1821, 1-9.
2015 Leung, J., Russell, B., Connell, S., Ng, J. & Lo, M. (2015). Acid dulls the senses: impaired locomotion and foraging performance in a marine mollusc. Animal Behaviour, 106, 223-229.
2015 Nagelkerken, I. & Connell, S. (2015). Global alteration of ocean ecosystem functioning due to increasing human CO₂ emissions. Proceedings of the National Academy of Sciences of the United States of America, 112, 43, 13272-13277.
2015 Pistevos, J., Nagelkerken, I., Rossi, T., Olmos, M. & Connell, S. (2015). Ocean acidification and global warming impair shark hunting behaviour and growth. Scientific Reports, 5, 1, 16293-1-16293-10.
2015 Gaylord, B., Kroeker, K., Sunday, J., Anderson, K., Barry, J., Brown, N. ... Harley, C. (2015). Ocean acidification through the lens of ecological theory. Ecology, 96, 1, 3-15.
2015 Alleway, H. & Connell, S. (2015). Loss of an ecological baseline through the eradication of oyster reefs from coastal ecosystems and human memory. Conservation Biology, 29, 3, 795-804.
2015 Kelaher, B., Tan, M., Figueira, W., Gillanders, B., Connell, S., Goldsworthy, S. ... Coleman, M. (2015). Fur seal activity moderates the effects of an Australian marine sanctuary on temperate reef fish. Biological Conservation, 182, 205-214.
2015 Connell, S. & Ghedini, G. (2015). Resisting regime-shifts: the stabilising effect of compensatory processes. Trends in Ecology & Evolution, 30, 9, 513-515.
2015 McSkimming, C., Tanner, J., Russell, B. & Connell, S. (2015). Compensation of nutrient pollution by herbivores in seagrass meadows. Journal of Experimental Marine Biology and Ecology, 471, 112-118.
2015 Ghedini, G., Russell, B. & Connell, S. (2015). Trophic compensation reinforces resistance: herbivory absorbs the increasing effects of multiple disturbances. Ecology Letters, 18, 2, 182-187.
2015 Hatton MacDonald, D., Ardeshiri, A., Rose, J., Russell, B. & Connell, S. (2015). Valuing coastal water quality: Adelaide, South Australia metropolitan area. Marine Policy, 52, 116-124.
2015 Ling, S., Scheibling, R., Rassweiler, A., Johnson, C., Shears, N., Connell, S. ... Johnson, L. (2015). Global regime shift dynamics of catastrophic sea urchin overgrazing. Philosophical Transactions of the Royal Society B: Biological Sciences, 370, 1659, 1-10.
2015 Falkenberg, L., Connell, S., Coffee, O., Ghedini, G. & Russell, B. (2015). Species interactions can maintain resistance of subtidal algal habitats to an increasingly modified world. Global Ecology and Conservation, 4, 549-558.
2015 Mertens, N., Russell, B. & Connell, S. (2015). Escaping herbivory: ocean warming as a refuge for primary producers where consumer metabolism and consumption cannot pursue. Oecologia, 179, 4, 1223-1229.
2014 Alleway, H. K., Connell, S. D., Ward, T. M. & Gillanders, B. M. (2014). Historical changes in mean trophic level of southern Australian fisheries. Marine and Freshwater Research, 65, 10, 884-893.
2014 Burnell, O., Russell, B., Irving, A. & Connell, S. (2014). Seagrass response to CO₂ contingent on epiphytic algae: indirect effects can overwhelm direct effects. Oecologia, 176, 3, 871-882.
2014 Gurgel, C., Wernberg, T., Thomsen, M., Russell, B., Adam, P., Waters, J. & Connell, S. (2014). Shared patterns of species turnover between seaweeds and seed plants break down at increasing distances from the sea. Ecology and Evolution, 4, 1, 27-34.
2014 Helmuth, B., Russell, B., Connell, S., Dong, Y., Harley, C., Lima, F. ... Mieszkowska, N. (2014). Beyond long-term averages: making biological sense of a rapidly changing world. Climate Change Responses, 1, 1, 6-1-6-12.
2014 Mohring, M. B., Wernberg, T., Wright, J. T., Connell, S. D. & Russell, B. (2014). Biogeographic variation in temperature drives performance of kelp gametophytes during warming. Marine Ecology Progress Series, 513, 85-96.
2014 Wernberg, T., Russell, B., Bradshaw, C., Gurgel, C., Thomsen, M., Poloczanska, E. & Connell, S. (2014). Misconceptions about analyses of Australian seaweed collections. Phycologia, 53, 3, 215-220.
2014 Smith, H., Anderson, M., Gillanders, B. & Connell, S. (2014). Longitudinal variation and effects of habitat on biodiversity of Australasian temperate reef fishes. Journal of Biogeography, 41, 11, 2128-2139.
2014 Burnell, O., Connell, S., Irving, A., Watling, J. & Russell, B. (2014). Contemporary reliance on bicarbonate acquisition predicts increased growth of seagrass Amphibolis antarctica in a high-CO₂ world. Conservation Physiology, 2, 1, cou052-1-cou052-11.
2014 Falkenberg, L., Connell, S. & Russell, B. (2014). Herbivory mediates the expansion of an algal habitat under nutrient and CO₂ enrichment. Marine Ecology Progress Series, 497, 87-92.
2014 Connell, S. D., Foster, M. S. & Airoldi, L. (2014). What are algal turfs? Towards a better description of turfs. Marine Ecology Progress Series, 495, 299-307.
2013 Burnell, O., Russell, B., Irving, A. & Connell, S. (2013). Eutrophication offsets increased sea urchin grazing on seagrass caused by ocean warming and acidification. Marine Ecology-Progress Series, 485, 37-46.
2013 Fordham, D., Mellin, C., Russell, B., Akcakaya, H., Bradshaw, C., Aiello-Lammens, M. ... Brook, B. (2013). Population dynamics can be more important than physiological limits for determining range shifts under climate change. Global Change Biology, 19, 10, 3224-3237.
2013 Falkenberg, L., Russell, B. & Connell, S. (2013). Future herbivory: the indirect effects of enriched CO₂ may rival its direct effects. Marine Ecology-Progress Series, 492, 85-95.
2013 Burnell, O., Connell, S., Irving, A. & Russell, B. (2013). Asymmetric patterns of recovery in two habitat forming seagrass species following simulated overgrazing by urchins. Journal of Experimental Marine Biology and Ecology, 448, 114-120.
2013 Russell, B., Connell, S., Findlay, H., Tait, K., Widdicombe, S. & Mieszkowska, N. (2013). Ocean acidification and rising temperatures may increase biofilm primary productivity but decrease grazer consumption. Philosophical Transactions of the Royal Society of London Series B-Biological Sciences, 368, 1627, 1-11.
2013 Ghedini, G., Russell, B. & Connell, S. (2013). Managing local coastal stressors to reduce the ecological effects of ocean acidification and warming. Water, 5, 4, 1653-1661.
2013 Connell, S., Kroeker, K., Fabricius, K., Kline, D. & Russell, B. (2013). The other ocean acidification problem: CO₂ as a resource among competitors for ecosystem dominance. Philosophical Transactions of the Royal Society of London Series B-Biological Sciences, 368, 1627, 1-10.
2013 Russell, B., Connell, S., Uthicke, S., Muehllehner, N., Fabricius, K. & Hall-Spencer, J. (2013). Future seagrass beds: can increased productivity lead to carbon storage?. Marine Pollution Bulletin, 73, 2, 463-469.
2013 Connell, S. (2013). By recognizing the broader landscape, we may better anticipate the outcome of local processes: editorial comment on the feature article by Miguel Matias. Marine Biology, 160, 3, 491-491.
2013 Falkenberg, L., Russell, B. & Connell, S. (2013). Contrasting resource limitations of marine primary producers: implications for competitive interactions under enriched CO₂ and nutrient regimes. Oecologia, 172, 2, 575-583.
2013 Wernberg, T., Thomsen, M., Connell, S., Russell, B., Waters, J., Zuccarello, G. ... Gurgel, C. (2013). The footprint of continental-scale ocean currents on the biogeography of seaweeds. PLoS One, 8, 11, 1-8.
2013 Falkenberg, L., Connell, S. & Russell, B. (2013). Disrupting the effects of synergies between stressors: improved water quality dampens the effects of future CO₂ on a marine habitat. Journal of Applied Ecology, 50, 1, 51-58.
2013 Coleman, M., Feng, M., Roughan, M., Cetina-Heredia, P. & Connell, S. (2013). Temperate shelf water dispersal by Australian boundary currents: implications for population connectivity. Limnology and Oceanography: Fluids and Environments, 3, 1, 295-309.
2012 Bulleri, F., Russell, B. & Connell, S. (2012). Context-dependency in the effects of nutrient loading and consumers on the availability of space in marine rocky environments. PLoS One, 7, 3, 1-9.
2012 Mellin, C., Russell, B., Connell, S., Brook, B. & Fordham, D. (2012). Geographic range determinants of two commercially important marine molluscs. Diversity and Distributions, 18, 2, 133-146.
2012 Russell, B., Harley, C., Wernberg, T., Mieszkowska, N., Widdicombe, S., Hall-Spencer, J. & Connell, S. (2012). Predicting ecosystem shifts requires new approaches that integrate the effects of climate change across entire systems. Biology Letters, 8, 2, 164-166.
2012 Russell, B. & Connell, S. (2012). Origins and consequences of global and local stressors: Incorporating climatic and non-climatic phenomena that buffer or accelerate ecological change. Marine Biology, 159, 11, 2633-2639.
2012 Livore, J. & Connell, S. (2012). Reducing per capita food supply alters urchin condition and habitat. Marine Biology, 159, 5, 967-973.
2012 Livore, J. & Connell, S. (2012). Fine-scale effects of sedentary urchins on canopy and understory algae. Journal of Experimental Marine Biology and Ecology, 411, 66-69.
2012 Livore, J. & Connell, S. (2012). Effects of food origin and availability on sea urchin condition and feeding behaviour. Journal of Sea Research, 68, 1-5.
2012 Russell, B., Connell, S., Mellin, C., Brook, B., Burnell, O. & Fordham, D. (2012). Predicting the distribution of commercially important invertebrate stocks under future climate. PLoS One, 7, 12, 1-9.
2012 Kline, D., Teneva, L., Schneider, K., Miard, T., Chai, A., Marker, M. ... Hoegh-Guldberg, O. (2012). A short-term in situ CO₂ enrichment experiment on Heron Island (GBR). Scientific Reports, 2, 1, 1-9.
2012 Falkenberg, L., Russell, B. & Connell, S. (2012). Stability of strong species interactions resist the synergistic effects of local and global pollution in kelp forests. PLoS One, 7, 3, 1-7.
2011 Irving, A., Connell, S. & Russell, B. (2011). Restoring coastal plants to improve global carbon storage: Reaping what we sow. PLoS One, 6, 3, 1-6.
2011 Wernberg-Moller, T., Russell, B., Thomsen, M., Gurgel, C., Bradshaw, C., Poloczanska, E. & Connell, S. (2011). Seaweed communities in retreat from ocean warming. Current Biology, 21, 21, 1828-1832.
2011 Coleman, M., Roughan, M., Macdonald, H., Connell, S., Gillanders, B., Kelaher, B. & Steinberg, P. (2011). Variation in the strength of continental boundary currents determines continent-wide connectivity in kelp. Journal of Ecology, 99, 4, 1026-1032.
2011 Connell, S., Russell, B. & Irving, A. (2011). Can strong consumer and producer effects be reconciled to better forecast 'catastrophic' phase-shifts in marine ecosystems?. Journal of Experimental Marine Biology and Ecology, 400, 1-2 Sp Iss, 296-301.
2011 Russell, B., Passarelli, C. & Connell, S. (2011). Forecasted CO₂ modifies the influence of light in shaping subtidal habitat. Journal of Phycology, 47, 4, 744-752.
2011 Watts, M., Li, Y., Russell, B., Mellin, C., Connell, S. & Fordham, D. (2011). A novel method for mapping reefs and subtidal rocky habitats using artificial neural networks. Ecological Modelling, 222, 15, 2606-2614.
2011 Wernberg, T., Russell, B., Moore, P., Ling, S., Smale, D., Campbell, A. ... Connell, S. (2011). Impacts of climate change in a global hotspot for temperate marine biodiversity and ocean warming. Journal of Experimental Marine Biology and Ecology, 400, 1-2 Sp Iss, 7-16.
2011 Colella, M., Parkinson, A., Evans, T., Robertson, J. & Roux, C. (2011). The Effect of Ionizing Gamma Radiation on Natural and Synthetic Fibers and Its Implications for the Forensic Examination of Fiber Evidence*. Journal of Forensic Sciences, 56, 3, 591-605.
2010 Falkenberg, L., Burnell, O., Connell, S. & Russell, B. (2010). Sustainability in Near-shore Marine Systems: Promoting Natural Resilience. Sustainability, Online, 8, 1-8.
2010 Coutts, A., Piola, R., Hewitt, C., Connell, S. & Gardner, J. (2010). Effect of vessel voyage speed on survival of biofouling organisms: implications for translocation of non-indigenous marine species. Biofouling, 26, 1, 1-13.
2010 Waters, J., Wernberg, T., Connell, S., Thomsen, M., Zuccarello, G., Kraft, G. ... Gurgel, C. (2010). Australia's marine biogeography revisited: Back to the future?. Austral Ecology, 35, 8, 988-992.
2010 Russell, B. (2010). Honing the geoengineering strategy. Science, 327, 5962, 144-145.
2010 Connell, S. & Russell, B. (2010). The direct effects of increasing CO₂ and temperature on non-calcifying organisms: increasing the potential for phase shifts in kelp forests. Proceedings of the Royal Society of London Series B-Biological Sciences, 277, 1686, 1409-1415.
2009 Coleman, M., Gillanders, B. & Connell, S. (2009). Dispersal and gene flow in the habitat-forming kelp, Ecklonia radiata: relative degrees of isolation across an east-west coastline. Marine and Freshwater Research, 60, 8, 802-809.
2009 Gorman, D., Russell, B. & Connell, S. (2009). Land-to-sea connectivity: Linking human-derived terrestrial subsidies to subtidal habitat change on open rocky coasts. Ecological Applications, 19, 5, 1114-1126.
2009 Elsdon, T. & Connell, S. (2009). Spatial and temporal monitoring of coastal water quality: refining the way we consider, gather, and interpret patterns. Aquatic Biology, 5, 2, 157-166.
2009 Saunders, T., Connell, S. & Mayfield, S. (2009). Differences in abalone growth and morphology between locations with high and low food availability: morphologically fixed or plastic traits?. Marine Biology, 156, 6, 1255-1263.
2009 Russell, B., Thompson, J. A., Falkenberg, L. & Connell, S. (2009). Synergistic effects of climate change and local stressors: CO₂ and nutrient-driven change in subtidal rocky habitats. Global Change Biology, 15, 9, 2153-2162.
2009 Gorman, D. & Connell, S. (2009). Recovering subtidal forests in human-dominated landscapes. Journal of Applied Ecology, 46, 6, 1258-1265.
2009 Russell, B. & Connell, S. (2009). Eutrophication science: moving into the future. Trends in Ecology & Evolution, 24, 10, 527-528.
2008 Ruesink, J. L. (2008). Marine Ecology . Edited by Sean D. Connell and , Bronwyn M. Gillanders. Oxford and New York: Oxford University Press. $75.00 (paper). xxxiii + 630 p.; ill.; index. 978‐0‐19‐555302‐4. 2007.. The Quarterly Review of Biology, 83, 2, 200-201.
2008 Wernberg-Moller, T. & Connell, S. (2008). Physical disturbance and subtidal habitat structure on open rocky coasts: Effects of wave exposure, extent and intensity. Journal of Sea Research, 59, 4, 237-248.
2008 Goodsell, P. & Connell, S. (2008). Complexity in the relationship between matrix composition and inter-patch distance in fragmented habitats. Marine Biology, 154, 1, 117-125.
2008 Connell, S., Russell, B., Turner, D., Shepherd, A., Kildea, T., Miller, D. ... Cheshire, A. (2008). Recovering a lost baseline: missing kelp forests from a metropolitan coast. Marine Ecology-Progress Series, 360, 63-72.
2008 Roberts, B. & Connell, S. (2008). Detecting benthic responses to human-induced change: effectiveness of alternate taxonomic classification and indices. Marine Ecology-Progress Series, 358, 75-84.
2008 Connell, S. & Irving, A. (2008). Integrating ecology with biogeography using landscape characteristics: a case study of subtidal habitat across continental Australia. Journal of Biogeography, 35, 9, 1608-1621.
2008 Valdivia, N., Stehbens, J., Hermelink, B., Connell, S., Molis, M. & Wahl, M. (2008). Disturbance mediates the effects of nutrients on developing assemblages of epibiota. Austral Ecology, 33, 8, 951-962.
2007 Glasby, T., Connell, S., Holloway, M. & Hewitt, C. (2007). Nonindigenous biota on artificial structures: could habitat creation facilitate biological invasions?. Marine Biology, 151, 3, 887-895.
2007 Russell, B. & Connell, S. (2007). Response of grazers to sudden nutrient pulses in oligotrophic versus eutrophic conditions. Marine Ecology-Progress Series, 349, 73-80.
2007 Fowler-Walker, M. & Connell, S. (2007). Habitat heterogeneity as a consequence of sub stratum-orientation and kelp-canopy: Relating interdependent responses to common patterns. Journal of Experimental Marine Biology and Ecology, 343, 1, 127-137.
2007 Coleman, M., Vytopil, E., Goodsell, P., Gillanders, B. & Connell, S. (2007). Diversity and depth-related patterns of mobile invertebrates associated with kelp forests. Marine and Freshwater Research, 58, 7, 589-595.
2007 Johnston, E., Connell, S., Irving, A., Pile, A. & Gillanders, B. (2007). Antarctic patterns of shallow subtidal habitat and inhabitants in Wilke' s Land. Polar Biology, 30, 6, 781-788.
2006 Fowler-Walker, M., Wernberg, T. & Connell, S. (2006). Differences in kelp morphology between wave sheltered and exposed localities: morphologically plastic or fixed traits?. Marine Biology, 148, 4, 755-767.
2006 Irving, A. & Connell, S. (2006). Physical disturbance by kelp abrades erect algae from the understorey. Marine Ecology-Progress Series, 324, 127-137.
2006 Irving, A. & Connell, S. (2006). Predicting understorey structure from the presence and composition of canopies: an assembly rule for marine algae. Oecologia, 148, 3, 491-502.
2005 Fowler-Walker, M., Gillanders, B., Connell, S. & Irving, A. (2005). Patterns of association between canopy-morphology and understorey assemblages across temperate Australia. Estuarine Coastal and Shelf Science, 63, 1-2, 133-141.
2005 Russell, B., Elsdon, T., Gillanders, B. & Connell, S. (2005). Nutrients increase epiphyte loads: broad-scale observations and an experimental assessment. Marine Biology, 147, 2, 551-558.
2005 Anderson, M., Connell, S., Gillanders, B., Diebel, C., Blom, W., Saunders, J. & Landers, T. (2005). Relationships between taxonomic resolution and spatial scales of multivariate variation. Journal of Animal Ecology, 74, 4, 636-646.
2005 Connell, S. (2005). Assembly and maintenance of subtidal habitat heterogeneity: synergistic effects of light penetration and sedimentation. Marine Ecology-Progress Series, 289, 53-61.
2005 Copertino, M., Connell, S. & Cheshire, A. (2005). The prevalence and production of turf-forming algae on a temperate subtidal coast. Phycologia, 44, 3, 241-248.
2005 Goodsell, P. & Connell, S. (2005). Disturbance initiates diversity in recruitment of canopy-forming algae: interactive effects of canopy-thinning and substratum availability. Phycologia, 44, 6, 632-639.
2005 Fowler-Walker, M., Connell, S. & Gillanders, B. (2005). To what extent do geographic and associated environmental variables correlate with kelp morphology across temperate Australia?. Marine and Freshwater Research, 56, 6, 877-887.
2005 Goodsell, P. & Connell, S. (2005). Historical configuration of habitat influences the effects of disturbance on mobile invertebrates. Marine Ecology-Progress Series, 299, 79-87.
2005 Irving, A., Connell, S., Johnston, E., Pile, A. & Gillanders, B. (2005). The response of encrusting coralline algae to canopy loss: an independent test of predictions on an Antarctic coast. Marine Biology, 147, 5, 1075-1083.
2005 Fowler-Walker, M., Connell, S. & Gillanders, B. (2005). Variation at local scales need not impede tests for broader scale patterns. Marine Biology, 147, 3, 823-831.
2005 Russell, B. & Connell, S. (2005). A novel interaction between nutrients and grazers alters relative dominance of marine habitats. Marine Ecology-Progress Series, 289, 5-11.
2005 Drummond, S. & Connell, S. (2005). Quantifying percentage cover of subtidal organisms on rocky coasts: a comparison of the costs and benefits of standard methods. Marine and Freshwater Research, 56, 6, 865-876.
2005 Russell, B., Gillanders, B. & Connell, S. (2005). Proximity and size of neighbouring habitat affects invertebrate diversity. Marine Ecology-Progress Series, 296, IS, 31-38.
2004 Goodsell, P., Fowler-Walker, M., Gillanders, B. & Connell, S. (2004). Variations in the configuration of algae in subtidal forests: Implications for invertebrate assemblages. Austral Ecology, 29, 3, 350-357.
2004 Gorgula, S. & Connell, S. (2004). Expansive covers of turf-forming algae on human-dominated coast: the relative effects of increasing nutrient and sediment loads. Marine Biology, 145, 3, 613-619.
2004 Irving, A., Connell, S. & Elsdon, T. (2004). Effects of kelp canopies on bleaching and photosynthetic activity of encrusting coralline algae. Journal of Experimental Marine Biology and Ecology, 310, 1, 1-12.
2004 Irving, A., Connell, S. & Gillanders, B. (2004). Local complexity in patterns of canopy-benthos associations produces regional patterns across temperate Australasia. Marine Biology, 144, 2, 361-368.
2003 Connell, S. (2003). The monopolization of understorey habitat by subtidal encrusting coralline algae: a test of the combined effects of canopy-mediated light and sedimentation. Marine Biology, 142, 6, 1065-1071.
2003 Connell, S. (2003). Effects of habitat amount vs habitat configuration on persistance of small vs large populations inhabiting patchy landscapes. Records of the South Australian Museum Monograph Series, 7, 61-66.
2003 Connell, S. (2003). Negative effects overpower the positive of kelp to exclude invertebrates from the understorey community. Oecologia, 137, 1, 97-103.
2003 Irving, A., Fowler-Walker, M. & Connell, S. (2003). Associations of forest-type with morphology of Ecklonia radiata and abundance of understorey algae. Transactions of the Royal Society of South Australia, 127, 2, 167-175.
2002 Connell, S. (2002). Effects of a predator and prey on a foraging reef fish: implications for understanding density-dependent growth. Journal of Fish Biology, 60, 6, 1551-1561.
2002 Irving, A. & Connell, S. (2002). Interactive effects of sedimentation and microtopography on the abundance of subtidal turf-forming algae. Phycologia, 41, 5, 517-522.
2002 Irving, A. & Connell, S. (2002). Sedimentation and light penetration interact to maintain heterogeneity of subtidal habitats: algal versus invertebrate dominated assemblages. Marine Ecology-Progress Series, 245, 83-91.
2002 Wellenreuther, M. & Connell, S. (2002). Response of predators to prey abundance: separating the effects of prey density and patch size. Journal of Experimental Marine Biology and Ecology, 273, 1, 61-71.
2002 Goodsell, P. & Connell, S. (2002). Can habitat loss be treated independently of habitat configuration? Implications for rare and common taxa in fragmented landscapes. Marine Ecology-Progress Series, 239, 37-44.
2002 Fowler-Walker, M. & Connell, S. (2002). Opposing states of subtidal habitat across temperate Australia: consistency and predictability in kelp canopy-benthic associations. Marine Ecology-Progress Series, 240, 49-56.
2002 Holloway, M. & Connell, S. (2002). Why do floating structures create novel habitats for subtidal epibiota?. Marine Ecology-Progress Series, 235, 43-52.
2001 Connell, S. (2001). Predatory fish do not always affect the early development of epibiotic assemblages. Journal of Experimental Marine Biology and Ecology, 260, 1, 1-12.
2001 Glasby, T. & Connell, S. (2001). Orientation and position of substrata have large effects on epibiotic assemblages. Marine Ecology-Progress Series, 214, 26 April, 127-135.
2001 Melville, A. & Connell, S. (2001). Experimental effects of kelp canopies on subtidal coralline algae. Austral Ecology, 26, 1, 102-108.
2001 Drabsch, S., Tanner, J. & Connell, S. (2001). Limited infaunal response to experimental trawling in previously untrawled areas. ICES Journal of Marine Science, 58, 6, 1261-1271.
2001 Coleman, M. & Connell, S. (2001). Weak effects of epibiota on the abundances of fishes associated with pier pilings in Sydney Harbour. Environmental Biology of Fishes, 61, 3, 231-239.
2001 Connell, S. (2001). Urban structures as marine habitats: an experimental comparison of the composition and abundance of subtidal epibiota among pilings, pontoons and rocky reefs. Marine Environmental Research, 52, 2, 115-125.
2001 Saunders, R. & Connell, S. (2001). Interactive effects of shade and surface orientation on the recruitment of spirorbid polychaetes. Austral Ecology, 26, 1, 109-115.
2000 Connell, S. (2000). Is there satety-in-numbers for prey?. Oikos, 88, 3, 527-532.
2000 Underwood, A., Chapman, M. & Connell, S. (2000). Observations in ecology: You can't make progress on processes without understanding the patterns. Journal of Experimental Marine Biology and Ecology, 250, 1-2, 97-115.
2000 Connell, S. (2000). Encounter rates of a juvenile reef fish with small and predatory fishes. S. Ross (Ed.). Copeia, 2000, 1, 36-41.
2000 Connell, S. (2000). Floating pontoons create novel habitats for subtidal epibiota. Journal of Experimental Marine Biology and Ecology, 247, 2, 183-194.
1999 Connell, S. & Lincoln-Smith, M. (1999). Depth and the structure of assemblages of demersal fish: experimental trawling along a temperate coast. Estuarine Coastal and Shelf Science, 48, 4, 483-495.
1999 Connell, S. & Glasby, T. (1999). Do urban structures influence local abundance and diversity of subtidal epibiota? A case study from Sydney Harbour, Australia. Marine Environmental Research, 47, 4, 373-387.
1999 Anderson, M. & Connell, S. (1999). Predation by fish on intertidal oysters. Marine Ecology Progress Series, 187, 203-211.
1999 Connell, S. & Anderson, M. (1999). Predation by fish on assemblages of intertidal epibiota: effects of predator size and patch size. Journal of Experimental Marine Biology and Ecology, 241, 1, 15-29.
1999 Glasby, T. & Connell, S. (1999). Urban structures as marine habitats. Ambio, 28, 7, 595-598.
1999 Connell, S. (1999). Effects of Surface Orientation on the Cover of Epibiota. Biofouling, 14, 3, 219-226.
1998 Connell, S. (1998). Patterns of pisciviory by resident predatory reef fish at One Tree Reef, Great Barrier Reef. Marine and Freshwater Research, 49, 1, 25-30.
1998 Connell, S. (1998). Effects of predators on growth, mortality and abundance of a juvenile reef-fish: Evidence from manipulations of predator and prey abundance. Marine Ecology Progress Series, 169, 251-261.
1998 Connell, S. & Kingsford, J. (1998). Spatial, temporal and habitat-related variation in the abundance of large predatory fish at One Tree Reef, Australia. Coral Reefs, 17, 1, 49-57.
1998 Connell, S., Samoilys, M., Lincoln Smith, M. & Leqata, J. (1998). Comparisons of abundance of coral-reef fish: Catch and effort surveys vs visual census. Austral Ecology, 23, 6, 579-586.
1997 Connell, S. (1997). Exclusion of predatory fish on a coral reef: The anticipation, pre- emption and evaluation of some caging artefacts. Journal of Experimental Marine Biology and Ecology, 213, 2, 181-198.
1997 Connell, S. (1997). The relationship between large predatory fish and recruitment and mortality of juvenile coral reef-fish on artificial reefs. Journal of Experimental Marine Biology and Ecology, 209, 1-2, 261-278.
1996 Connell, S. (1996). Variations in mortality of a coral reef fish: Links with predator abundance. Marine Biology, 126, 2, 347-352.
1991 Connell, S. & Jones, G. (1991). The influence of habitat complexity on postrecruitment processes in a temperate reef fish population. Journal of Experimental Marine Biology and Ecology, 151, 2, 271-294.


Year Citation
2007 S. Connell & B. Gillanders (Eds.) (2007). Marine ecology. South Melbourne: Oxford University Press.

Book Chapters

Year Citation
2014 Wernberg, T., Russell, B., Thomsen, M. S. & Connell, S. D. (2014). Marine biodiversity and climate change. In B. Freedman (Ed.), Global Environmental Change (pp. 181-187). Springer.
2014 Russell, B. D. & Connell, S. D. (2014). Marine benthic productivity. In Y. Wang (Ed.), Encyclopedia of Natural Resources: Water (pp. 767-771). New York: Taylor and Francis.
2014 Russell, B. & Connell, S. (2014). Ecosystem resilience and resistance to climate change. In B. Freedman (Ed.), Global Environmental Change (pp. 133-139). Springer.
2013 Falkenberg, L., Connell, S. & Russell, B. (2013). Current and future eutrophication of nearshore marine environments: Causes, consequences and potential management strategies. In Eutrophication: Causes, Economic Implications and Future Challenges (pp. 215-238).
2009 Harley, C. & Connell, S. (2009). Shifts in abiotic variables and consequences for diversity. In Martin Wahl (Ed.), Marine hard bottom communities (pp. 257-268). Heidelberg, Germany: Springer.
2009 Airoldi, L., Connell, S. & Beck, M. (2009). The loss of natural habitats and the addition of artificial substrata. In Martin Wahl (Ed.), Marine hard bottom communities (pp. 269-280). Heidelberg, Germany: Springer.
2009 Connell, S. & Irving, A. (2009). The subtidal ecology of rocky coasts: Local-Regional-Biogeographic patterns and their experimental analysis. In J. D. Witman & K. Roy (Eds.), Marine Macroecology (pp. 392-418). USA: University of Chicago Press.
2007 Connell, S. (2007). Water quality and the loss of coral reefs and kelp forests: alternative states and the influence of fishing. In Marine ecology (pp. 556-568). South Melbourne: Oxford University Press.
2007 Connell, S. & Vanderklift, M. (2007). Negative interactions: The influence of predators and herbivores on prey and ecological systems. In Marine ecology (pp. 72-100). South Melbourne: Oxford University Press.
2007 Connell, S. (2007). Subtidal temperate rocky habitats: habitat heterogenity at local to continental scales. In Marine ecology (pp. 378-401). South Melbourne: Oxford University Press.
2005 Connell, S. & Gillanders, B. (2005). Urban Ecological Communities 4: The marine and coastal environment. In Adelaide - Nature of a city: The Ecology of a Dynamic City from 1836 to 2036 (pp. 298-318). Adelaide: BioCity: Centre for Urban Habitats.

Conference Papers

Year Citation
2016 McSkimming, C., Russell, B., Tanner, J. & Connell, S. (2016). A test of metabolic and consumptive responses to local and global perturbations: enhanced resources stimulate herbivores to counter expansion of weedy species. 10th International Temperate Reefs Symposium. Perth, W.A..
2016 Bennett, S., Wernberg, T., Connell, S., Hobday, A., Johnson, C. & Poloczanska, E. (2016). The 'Great Southern Reef': social, ecological and economic value of Australia's neglected kelp forests. 10th International Temperate Reefs Symposium. Perth, W.A..
2016 Alleway, H., Thurstan, R., Lauer, P. & Connell, S. (2016). Incorporating historical data into aquaculture planning. 3rd International Symposium on the Effects of Climate Change on the World's Oceans. PICES, Santos, BRAZIL.


   Me writing a grant @ in the middle of the Great Australian Bight.

   My best ideas come whilst surrounded by the systems I study.




Grants with layperson titles:

  • ARC Discovery (2014-2016):  Ocean acidification propagates resource enrichment and its intensifying consumption in the web of life.
  • ARC Future Fellowship (2011-2013):  Global and local emissions boosts limiting resources for bottom-up global change.
  • ARC Linkage (2008 -2010): Managing global imprints with local activities to empower local managers.
  • ARC Linkage (2005-2007): Planning for coastal policy success by resolving land-to-sea linkages.
  • ARC Queen Elizabeth Fellowship II (2003-2005): Ecosystem decline in human-dominated landscapes.
  • ARC Discovery (2002-2005): Biogeography and ecology of southern kelp forests.
  • ARC Discovery (2000-2003): Experimental ecology of a subtidal rocky coast.






I teach the hard skills with love and passion.

1. Marine Biology.  I coordinate the BSc (Marine Biology).  We have taken students to China and East Timor on fully paid trips.

2. Statistics.  I coordinate the 3rd Year statistics course for biologists.  I receive unsolicited emails, such as "I just wanted to thank you for being a highly effective and thoughtful teacher during the first four weeks of RMEB III this semester.  I feel confident for the remainder of the course because of the way you presented the (relatively dry!) content and the way you listened to the students' needs"  (Student, 2017).

3. Writing.  Beautiful and nourishing reading comes from the effort we put into writing. Relative to maths, this one of the most under-valued skills in science.  Science writing is so dull and impenetrable that it sounds as if we are speaking in tongues.  Science communicators are employed to decode the babble.  I teach how to write clear and simple - the way to the truth - with a twist for a lively read.




I have graduated over 100 research students.  I work with students to design a project they can be passionate about, with an focus on the spirit of discovery and the needs of society.  My students have careers as marine biologists:

  1. Universities (national and international)
  2. Departments of conservation and wildlife
  3. Government water utilities
  4. Private consultancies
  5. Fisheries institutes
  6. Local councils
  7. Marine Parks






Date Role Membership Country
2013 - ongoing Member The Environment Institute Australia

Committee Memberships

Date Role Committee Institution Country
2015 - ongoing Member Habitat Enhancement Working Group Primary Industries South Australia Australia
2014 - 2016 Member International Temperate Reefs Symposium (Plymouth, UK) Plymouth University United Kingdom
2008 - 2016 Chair International Temperate Reefs Sympoium (Adelaide)
2002 - 2014 Advisory Board Member Scientific Working Group Department of Environment, Water and Natural Resources Australia
2001 - 2008 Board Member Wildlife Advsory Group Department of Environment and Hertiage Australia

Editorial Boards

Date Role Editorial Board Name Country
2010 - 2013 Associate Editor Aquatic Biology
2004 - ongoing Associate Editor Marine Biology: international journal on life in oceans and coastal waters

Industry Partnerships

Date Engagement Type Partner Name
2013 - ongoing Consultant SA Water


Date Institution Department Organisation Type Country
2006 - 2012 Department of Agriculture Great Australian Bight Fishery Australia
1994 - 1996 The Ecology Lab Australia
8313 4364
North Terrace
Darling Building
Room Number
1 17
Org Unit
Ecology and Environmental Science