Professor Sean Connell

Sean Connell
School of Biological Sciences
Faculty of Sciences

Sea creatures, their life and their future - is my work. This daily life has inspired new policy & cleaner coasts by scientific discovery & saner theory.

Join me. I'm looking for students interested in biology and marine biology (PhD, Honours & MSc students). All projects are possible. Those underway include climate change and restoration ecology (e.g. restoring 20 hectares of oyster reef). Students snorkel & dive or use mesocosms & aquariums. These techniques enable a variety of projects - from behaviour through tests of how individuals work together to form communities.

Professor Sean Connell

Sea creatures, their life and their future - is my work. This daily life has inspired new policy & cleaner coasts by scientific discovery & saner theory.

Join me. I'm looking for students interested in biology and marine biology (PhD, Honours & MSc students). All projects are possible. Those underway include climate change and restoration ecology (e.g. restoring 20 hectares of oyster reef). Students snorkel & dive or use mesocosms & aquariums. These techniques enable a variety of projects - from behaviour through tests of how individuals work together to form communities.

Add these ingredients (below); work, people, fun, adventure and data...

Sean @ Work
       Sean @ Work
  With strangers
           Having fun
        In exotic locations
Sean writing down hard won data
    Getting data

... to get results (below)...

Collecting data in exotic locations with strangers who are fun to work with.  For students, visit my personal website and supervision tab on this page.

a a Book
a a a







Date Position Institution name
2011 Professor University of Adelaide
2005 - 2010 Associate Professor University of Adelaide
1999 - 2004 Senior Lecturer The University of Adelaide
1997 - 1999 Lecturer University of Sydney
1996 - 1997 Lecturer University of Tasmania (Australian Maritime College)
1994 - 1996 Scientist The Ecology Laboratory

Awards and Achievements

Date Type Title Institution Name Amount
2010 ARC Future Fellowship
2004 ARC Queen Elizabeath II Fellowship
2003 J.G. Russell Award Australian Academy of Science
2003 The Andrewartha Medal
1992 Scholarship (PhD) - Australia University of Sydney
1990 Scholarship (PhD) - New Zealand New Zealand University Grants Committee


Date Institution name Country Title
University of Sydney Australia PhD
University of Canterbury New Zealand BSc
University of Auckland New Zealand MSc (1st Class Honours)


Aquatic Ecosystem Studies and Stock Assessment, Climate Change Processes, Community Ecology, Ecological Impacts of Climate Change, Ecology, Ecosystem Function, Marine and Estuarine Ecology


Date Citation
2017 Harley,C, Connell,S, Doubleday,Z, Kelaher,B, Russell,B, Sarà,G, Helmuth,B, 2017, Conceptualizing ecosystem tipping points within a physiological framework, Ecology and Evolution, - 10.1002/ece3.3164
2017 Goldenberg,S, Nagelkerken,I, Ferreira,C, Ullah,H, Connell,S, 2017, Boosted food web productivity through ocean acidification collapses under warming, Global Change Biology, Online, 1-8 10.1111/gcb.13699
2017 Connell,SD, Fernandes,M, Burnell,OW, Doubleday,ZA, Griffin,KJ, Irving,AD, Leung,JY, Owen,S, Russell,BD, Falkenberg,LJ et al, 2017, Testing for thresholds of ecosystem collapse in seagrass meadows., Conservation biology : the journal of the Society for Conservation Biology, - 10.1111/cobi.12951
2017 Connell,S, Doubleday,Z, Hamlyn,S, Foster,N, Harley,C, Helmuth,B, Kelaher,B, Nagelkerken,I, Sarà,G, Russell,B et al, 2017, How ocean acidification can benefit calcifiers, Current Biology, 27, 3, R95-R96 10.1016/j.cub.2016.12.004
2017 Rossi,T, Connell,S, Nagelkerken,I, 2017, The sounds of silence: regime shifts impoverish marine soundscapes, Landscape Ecology, 32, 2, 239-248 10.1007/s10980-016-0439-x
2017 Pistevos,J, Nagelkerken,I, Rossi,T, Connell,S, 2017, Antagonistic effects of ocean acidification and warming on hunting sharks, Oikos, 126, 2, 241-247 10.1111/oik.03182
2017 Pistevos,J, Nagelkerken,I, Rossi,T, Connell,S, 2017, Ocean acidification alters temperature and salinity preferences in larval fish, Oecologia, 183, 2, 545-553 10.1007/s00442-016-3778-z
2017 Ghedini,G, Connell,S, 2017, Moving ocean acidification research beyond a simple science: Investigating ecological change and their stabilizers, Food Webs, Online, 1-7 10.1016/j.fooweb.2017.03.003
2017 Sunday,J, Fabricius,K, Kroeker,K, Anderson,K, Brown,N, Barry,J, Connell,S, Dupont,S, Gaylord,B, Hall-Spencer,J et al, 2017, Ocean acidification can mediate biodiversity shifts by changing biogenic habitat, Nature Climate Change, 7, 1, 81-85 10.1038/nclimate3161
2017 Nagelkerken,I, Goldenberg,S, Ferreira,C, Russell,B, Connell,S, 2017, Species Interactions Drive Fish Biodiversity Loss in a High-CO2 World, Current Biology, - 10.1016/j.cub.2017.06.023
2017 Provost,E, Kelaher,B, Dworjanyn,S, Russell,B, Connell,S, Ghedini,G, Gillanders,B, Figueira,W, Coleman,M, 2017, Climate-driven disparities among ecological interactions threaten kelp forest persistence, Global Change Biology, 23, 1, 353-361 10.1111/gcb.13414
2017 Hardy,N, Berry,T, Kelaher,B, Goldsworthy,S, Bunce,M, Coleman,M, Gillanders,B, Connell,S, Blewitt,M, Figueira,W et al, 2017, Assessing the trophic ecology of top predators across a recolonisation frontier using DNA metabarcoding of diets, Marine Ecology Progress Series, 573, 237-254 10.3354/meps12165
2017 Leung,J, Russell,B, Connell,S, 2017, Mineralogical plasticity acts as a compensatory mechanism to the impacts of ocean acidification, Environmental Science and Technology, 51, 5, 2652-2659 10.1021/acs.est.6b04709
2016 McSkimming,C, Connell,SD, Russell,BD, Tanner,JE, 2016, Habitat restoration: early signs and extent of faunal recovery relative to seagrass recovery, Estuarine, Coastal and Shelf Science, 171, 51-57 10.1016/j.ecss.2016.01.028
2016 Alleway,H, Gillanders,B, Connell,S, 2016, 'Neo-Europe' and its ecological consequences: the example of systematic degradation in Australia's inland fisheries, BIOLOGY LETTERS, 12, 1, 1-7 10.1098/rsbl.2015.0774
2016 Falkenberg,L, Russell,B, Connell,S, 2016, Design and performance evaluation of a mesocosm facility and techniques to simulate ocean acidification and warming, Limnology and Oceanography: Methods, 14, 4, 278-291 10.1002/lom3.10088
2016 Rossi,T, Nagelkerken,I, Pistevos,J, Connell,S, 2016, Lost at sea: ocean acidification undermines larval fish orientation via altered hearing and marine soundscape modification, Biology Letters, 12, 1, 1-4 10.1098/rsbl.2015.0937
2016 Ghedini,G, Connell,S, 2016, Organismal homeostasis buffers the effects of abiotic change on community dynamics, Ecology, 97, 10, 2671-2679 10.1002/ecy.1488
2016 Heldt,K, Connell,S, Anderson,K, Russell,B, Munguia,P, 2016, Future climate stimulates population out-breaks by relaxing constraints on reproduction, Scientific Reports, 6, 1, 33383-1-33383-8 10.1038/srep33383
2016 Nagelkerken,I, Russell,B, Gillanders,B, Connell,S, 2016, Ocean acidification alters fish populations indirectly through habitat modification, Nature Climate Change, 6, 1, 89-93 10.1038/nclimate2757
2016 Krumhansl,K, Okamoto,D, Rassweiler,A, Novak,M, Bolton,J, Cavanaugh,K, Connell,S, Johnson,C, Konar,B, Ling,S et al, 2016, Global patterns of kelp forest change over the past half-century, Proceedings of the National Academy of Sciences of the United States of America, 113, 48, 13785-13790 10.1073/pnas.1606102113
2016 Connell,S, Nimmo,D, Ghedini,G, Mac Nally,R, Bennett,A, 2016, Ecological Resistance - Why Mechanisms Matter: A Reply to Sundstrom et al., Trends in Ecology and Evolution, 31, 6, 413-414 10.1016/j.tree.2016.03.015
2016 Rossi,T, Connell,S, Nagelkerken,I, 2016, Silent oceans: ocean acidification impoverishes natural soundscapes by altering sound production of the world’s noisiest marine invertebrate, Proceedings of the Royal Society B, 283, 1826, 20153046-1-20153046-7 10.1098/rspb.2015.3046
2015 Gaylord,B, Kroeker,K, Sunday,J, Anderson,K, Barry,J, Brown,N, Connell,S, Dupont,S, Fabricius,K, Hall-Spencer,J et al, 2015, Ocean acidification through the lens of ecological theory, Ecology, 96, 1, 3-15 10.1890/14-0802.1
2015 Pistevos,J, Nagelkerken,I, Rossi,T, Olmos,M, Connell,S, 2015, Ocean acidification and global warming impair shark hunting behaviour and growth, Scientific Reports, 5, 1, 16293-1-16293-10 10.1038/srep16293
2015 Connell,S, Ghedini,G, 2015, Resisting regime-shifts: the stabilising effect of compensatory processes, Trends in Ecology & Evolution, 30, 9, 513-515 10.1016/j.tree.2015.06.014
2015 Hatton MacDonald,D, Ardeshiri,A, Rose,J, Russell,B, Connell,S, 2015, Valuing coastal water quality: Adelaide, South Australia metropolitan area, Marine Policy, 52, 116-124 10.1016/j.marpol.2014.11.003
2015 Kelaher,B, Tan,M, Figueira,W, Gillanders,B, Connell,S, Goldsworthy,S, Hardy,N, Coleman,M, 2015, Fur seal activity moderates the effects of an Australian marine sanctuary on temperate reef fish, Biological Conservation, 182, 205-214 10.1016/j.biocon.2014.12.011
2015 Ling,S, Scheibling,R, Rassweiler,A, Johnson,C, Shears,N, Connell,S, Salomon,A, Norderhaug,K, Pérez-Matus,A, Hernández,J et al, 2015, Global regime shift dynamics of catastrophic sea urchin overgrazing, Philosophical Transactions of the Royal Society B: Biological Sciences, 370, 1659, 1-10 10.1098/rstb.2013.0269
2015 Leung,J, Russell,B, Connell,S, Ng,J, Lo,M, 2015, Acid dulls the senses: impaired locomotion and foraging performance in a marine mollusc, Animal Behaviour, 106, 223-229 10.1016/j.anbehav.2015.06.004
2015 Mertens,N, Russell,B, Connell,S, 2015, Escaping herbivory: ocean warming as a refuge for primary producers where consumer metabolism and consumption cannot pursue, Oecologia, 179, 4, 1223-1229 10.1007/s00442-015-3438-8
2015 Alleway,H, Connell,S, 2015, Loss of an ecological baseline through the eradication of oyster reefs from coastal ecosystems and human memory, Conservation Biology, 29, 3, 795-804 10.1111/cobi.12452
2015 McSkimming,C, Tanner,J, Russell,B, Connell,S, 2015, Compensation of nutrient pollution by herbivores in seagrass meadows, Journal of Experimental Marine Biology and Ecology, 471, 112-118 10.1016/j.jembe.2015.05.018
2015 Nagelkerken,I, Connell,S, 2015, Global alteration of ocean ecosystem functioning due to increasing human CO₂ emissions, Proceedings of the National Academy of Sciences of the United States of America, 112, 43, 13272-13277 10.1073/pnas.1510856112
2015 Ghedini,G, Russell,B, Connell,S, 2015, Trophic compensation reinforces resistance: herbivory absorbs the increasing effects of multiple disturbances, Ecology Letters, 18, 2, 182-187 10.1111/ele.12405
2015 Falkenberg,L, Connell,S, Coffee,O, Ghedini,G, Russell,B, 2015, Species interactions can maintain resistance of subtidal algal habitats to an increasingly modified world, Global Ecology and Conservation, 4, 549-558 10.1016/j.gecco.2015.10.003
2015 Rossi,T, Nagelkerken,I, Simpson,S, Pistevos,J, Watson,S, Merillet,L, Fraser,P, Munday,P, Connell,S, 2015, Ocean acidification boosts larval fish development but reduces the window of opportunity for successful settlement, Proceedings of the Royal Society B, 282, 1821, 1-9 10.1098/rspb.2015.1954
2014 Burnell,O, Connell,S, Irving,A, Watling,J, Russell,B, 2014, Contemporary reliance on bicarbonate acquisition predicts increased growth of seagrass Amphibolis antarctica in a high-CO₂ world, Conservation Physiology, 2, 1, cou052-1-cou052-11 10.1093/conphys/cou052
2014 Connell,SD, Foster,MS, Airoldi,L, 2014, What are algal turfs? Towards a better description of turfs, Marine Ecology Progress Series, 495, 299-307 10.3354/meps10513
2014 Burnell,O, Russell,B, Irving,A, Connell,S, 2014, Seagrass response to CO₂ contingent on epiphytic algae: indirect effects can overwhelm direct effects, Oecologia, 176, 3, 871-882 10.1007/s00442-014-3054-z
2014 Smith,H, Anderson,M, Gillanders,B, Connell,S, 2014, Longitudinal variation and effects of habitat on biodiversity of Australasian temperate reef fishes, Journal of Biogeography, 41, 11, 2128-2139 10.1111/jbi.12359
2014 Wernberg,T, Russell,B, Bradshaw,C, Gurgel,C, Thomsen,M, Poloczanska,E, Connell,S, 2014, Misconceptions about analyses of Australian seaweed collections, Phycologia, 53, 3, 215-220 10.2216/13-197.1
2014 Mohring,MB, Wernberg,T, Wright,JT, Connell,SD, Russell,B, 2014, Biogeographic variation in temperature drives performance of kelp gametophytes during warming, Marine Ecology Progress Series, 513, 85-96 10.3354/meps10916
2014 Falkenberg,L, Connell,S, Russell,B, 2014, Herbivory mediates the expansion of an algal habitat under nutrient and CO₂ enrichment, Marine Ecology Progress Series, 497, 87-92 10.3354/meps10557
2014 Alleway,HK, Connell,SD, Ward,TM, Gillanders,BM, 2014, Historical changes in mean trophic level of southern Australian fisheries, Marine and Freshwater Research, 65, 10, 884-893 10.1071/MF13246
2014 Gurgel,C, Wernberg,T, Thomsen,M, Russell,B, Adam,P, Waters,J, Connell,S, 2014, Shared patterns of species turnover between seaweeds and seed plants break down at increasing distances from the sea, Ecology and Evolution, 4, 1, 27-34 10.1002/ece3.893
2014 Helmuth,B, Russell,B, Connell,S, Dong,Y, Harley,C, Lima,F, Sará,G, Williams,G, Mieszkowska,N, 2014, Beyond long-term averages: making biological sense of a rapidly changing world, Climate Change Responses, 1, 1, 6-1-6-12 10.1186/s40665-014-0006-0
2013 Russell,B, Connell,S, Uthicke,S, Muehllehner,N, Fabricius,K, Hall-Spencer,J, 2013, Future seagrass beds: can increased productivity lead to carbon storage?, Marine Pollution Bulletin, 73, 2, 463-469 10.1016/j.marpolbul.2013.01.031
2013 Falkenberg,L, Russell,B, Connell,S, 2013, Contrasting resource limitations of marine primary producers: implications for competitive interactions under enriched CO₂ and nutrient regimes, Oecologia, 172, 2, 575-583 10.1007/s00442-012-2507-5
2013 Ghedini,G, Russell,B, Connell,S, 2013, Managing local coastal stressors to reduce the ecological effects of ocean acidification and warming, Water, 5, 4, 1653-1661 10.3390/w5041653
2013 Russell,B, Connell,S, Findlay,H, Tait,K, Widdicombe,S, Mieszkowska,N, 2013, Ocean acidification and rising temperatures may increase biofilm primary productivity but decrease grazer consumption, Philosophical Transactions of the Royal Society of London Series B-Biological Sciences, 368, 1627, 1-11 10.1098/rstb.2012.0438
2013 Burnell,O, Russell,B, Irving,A, Connell,S, 2013, Eutrophication offsets increased sea urchin grazing on seagrass caused by ocean warming and acidification, Marine Ecology-Progress Series, 485, 37-46 10.3354/meps10323
2013 Fordham,D, Mellin,C, Russell,B, Akcakaya,H, Bradshaw,C, Aiello-Lammens,M, Caley,M, Connell,S, Mayfield,S, Shepherd,S et al, 2013, Population dynamics can be more important than physiological limits for determining range shifts under climate change, Global Change Biology, 19, 10, 3224-3237 10.1111/gcb.12289
2013 Wernberg,T, Thomsen,M, Connell,S, Russell,B, Waters,J, Zuccarello,G, Kraft,G, Sanderson,J, West,J, Gurgel,C et al, 2013, The footprint of continental-scale ocean currents on the biogeography of seaweeds, PLoS One, 8, 11, 1-8 10.1371/journal.pone.0080168
2013 Connell,S, 2013, By recognizing the broader landscape, we may better anticipate the outcome of local processes: editorial comment on the feature article by Miguel Matias, Marine Biology, 160, 3, 491-491 10.1007/s00227-012-2161-9
2013 Burnell,O, Connell,S, Irving,A, Russell,B, 2013, Asymmetric patterns of recovery in two habitat forming seagrass species following simulated overgrazing by urchins, Journal of Experimental Marine Biology and Ecology, 448, 114-120 10.1016/j.jembe.2013.07.003
2013 Connell,S, Kroeker,K, Fabricius,K, Kline,D, Russell,B, 2013, The other ocean acidification problem: CO₂ as a resource among competitors for ecosystem dominance, Philosophical Transactions of the Royal Society of London Series B-Biological Sciences, 368, 1627, 1-10 10.1098/rstb.2012.0442
2013 Falkenberg,L, Connell,S, Russell,B, 2013, Disrupting the effects of synergies between stressors: improved water quality dampens the effects of future CO₂ on a marine habitat, Journal of Applied Ecology, 50, 1, 51-58 10.1111/1365-2664.12019
2013 Falkenberg,L, Russell,B, Connell,S, 2013, Future herbivory: the indirect effects of enriched CO₂ may rival its direct effects, Marine Ecology-Progress Series, 492, 85-95 10.3354/meps10491
2013 Coleman,M, Feng,M, Roughan,M, Cetina-Heredia,P, Connell,S, 2013, Temperate shelf water dispersal by Australian boundary currents: implications for population connectivity, Limnology and Oceanography: Fluids and Environments, 3, 1, 295-309 10.1215/21573689-2409306
2012 Kline,D, Teneva,L, Schneider,K, Miard,T, Chai,A, Marker,M, Headley,K, Opdyke,B, Nash,M, Valetich,M et al, 2012, A short-term in situ CO₂ enrichment experiment on Heron Island (GBR), Scientific Reports, 2, 1-9 10.1038/srep00413
2012 Livore,J, Connell,S, 2012, Reducing per capita food supply alters urchin condition and habitat, Marine Biology, 159, 5, 967-973 10.1007/s00227-011-1875-4
2012 Russell,B, Connell,S, Mellin,C, Brook,B, Burnell,O, Fordham,D, 2012, Predicting the distribution of commercially important invertebrate stocks under future climate, PLoS One, 7, 12, 1-9 10.1371/journal.pone.0046554
2012 Russell,B, Harley,C, Wernberg,T, Mieszkowska,N, Widdicombe,S, Hall-Spencer,J, Connell,S, 2012, Predicting ecosystem shifts requires new approaches that integrate the effects of climate change across entire systems, Biology Letters, 8, 2, 164-166 10.1098/rsbl.2011.0779
2012 Livore,J, Connell,S, 2012, Fine-scale effects of sedentary urchins on canopy and understory algae, Journal of Experimental Marine Biology and Ecology, 411, 66-69 10.1016/j.jembe.2011.11.001
2012 Livore,J, Connell,S, 2012, Effects of food origin and availability on sea urchin condition and feeding behaviour, Journal of Sea Research, 68, 1-5 10.1016/j.seares.2011.11.002
2012 Russell,B, Connell,S, 2012, Origins and consequences of global and local stressors: Incorporating climatic and non-climatic phenomena that buffer or accelerate ecological change, Marine Biology, 159, 11, 2633-2639 10.1007/s00227-011-1863-8
2012 Falkenberg,L, Russell,B, Connell,S, 2012, Stability of strong species interactions resist the synergistic effects of local and global pollution in kelp forests, PLoS One, 7, 3, 1-7 10.1371/journal.pone.0033841
2012 Bulleri,F, Russell,B, Connell,S, 2012, Context-dependency in the effects of nutrient loading and consumers on the availability of space in marine rocky environments, PLoS One, 7, 3, 1-9 10.1371/journal.pone.0033825
2012 Mellin,C, Russell,B, Connell,S, Brook,B, Fordham,D, 2012, Geographic range determinants of two commercially important marine molluscs, Diversity and Distributions, 18, 2, 133-146 10.1111/j.1472-4642.2011.00822.x
2011 Watts,M, Li,Y, Russell,B, Mellin,C, Connell,S, Fordham,D, 2011, A novel method for mapping reefs and subtidal rocky habitats using artificial neural networks, Ecological Modelling, 222, 15, 2606-2614 10.1016/j.ecolmodel.2011.04.024
2011 Wernberg,T, Russell,B, Moore,P, Ling,S, Smale,D, Campbell,A, Coleman,M, Steinberg,P, Kendrick,G, Connell,S et al, 2011, Impacts of climate change in a global hotspot for temperate marine biodiversity and ocean warming, Journal of Experimental Marine Biology and Ecology, 400, 1-2 Sp Iss, 7-16 10.1016/j.jembe.2011.02.021
2011 Russell,B, Passarelli,C, Connell,S, 2011, Forecasted CO₂ modifies the influence of light in shaping subtidal habitat, Journal of Phycology, 47, 4, 744-752 10.1111/j.1529-8817.2011.01002.x
2011 Wernberg-Moller,T, Russell,B, Thomsen,M, Gurgel,C, Bradshaw,C, Poloczanska,E, Connell,S, 2011, Seaweed communities in retreat from ocean warming, Current Biology, 21, 21, 1828-1832 10.1016/j.cub.2011.09.028
2011 Irving,A, Connell,S, Russell,B, 2011, Restoring coastal plants to improve global carbon storage: Reaping what we sow, PLoS One, 6, 3, 1-6 10.1371/journal.pone.0018311
2011 Connell,S, Russell,B, Irving,A, 2011, Can strong consumer and producer effects be reconciled to better forecast 'catastrophic' phase-shifts in marine ecosystems?, Journal of Experimental Marine Biology and Ecology, 400, 1-2 Sp Iss, 296-301 10.1016/j.jembe.2011.02.031
2011 Coleman,M, Roughan,M, Macdonald,H, Connell,S, Gillanders,B, Kelaher,B, Steinberg,P, 2011, Variation in the strength of continental boundary currents determines continent-wide connectivity in kelp, Journal of Ecology, 99, 4, 1026-1032 10.1111/j.1365-2745.2011.01822.x
2010 Coutts,A, Piola,R, Hewitt,C, Connell,S, Gardner,J, 2010, Effect of vessel voyage speed on survival of biofouling organisms: implications for translocation of non-indigenous marine species, Biofouling, 26, 1, 1-13 10.1080/08927010903174599
2010 Russell,B, 2010, Honing the geoengineering strategy, Science, 327, 5962, 144-145 10.1126/science.327.5962.144-c
2010 Connell,S, Russell,B, 2010, The direct effects of increasing CO₂ and temperature on non-calcifying organisms: increasing the potential for phase shifts in kelp forests, Proceedings of the Royal Society of London Series B-Biological Sciences, 277, 1686, 1409-1415 10.1098/rspb.2009.2069
2010 Waters,J, Wernberg,T, Connell,S, Thomsen,M, Zuccarello,G, Kraft,G, Sanderson,J, West,J, Gurgel,C, 2010, Australia's marine biogeography revisited: Back to the future?, Austral Ecology, 35, 8, 988-992 10.1111/j.1442-9993.2010.02114.x
2010 Falkenberg,L, Burnell,O, Connell,S, Russell,B, 2010, Sustainability in Near-shore Marine Systems: Promoting Natural Resilience, Sustainability, Online, 8, 1-8 10.3390/su2082593
2009 Saunders,T, Connell,S, Mayfield,S, 2009, Differences in abalone growth and morphology between locations with high and low food availability: morphologically fixed or plastic traits?, Marine Biology, 156, 6, 1255-1263 10.1007/s00227-009-1167-4
2009 Gorman,D, Connell,S, 2009, Recovering subtidal forests in human-dominated landscapes, Journal of Applied Ecology, 46, 6, 1258-1265 10.1111/j.1365-2664.2009.01711.x
2009 Elsdon,T, Connell,S, 2009, Spatial and temporal monitoring of coastal water quality: refining the way we consider, gather, and interpret patterns, Aquatic Biology, 5, 2, 157-166 10.3354/ab00146
2009 Russell,B, Thompson,JA, Falkenberg,L, Connell,S, 2009, Synergistic effects of climate change and local stressors: CO₂ and nutrient-driven change in subtidal rocky habitats, Global Change Biology, 15, 9, 2153-2162 10.1111/j.1365-2486.2009.01886.x
2009 Russell,B, Connell,S, 2009, Eutrophication science: moving into the future, Trends in Ecology & Evolution, 24, 10, 527-528 10.1016/j.tree.2009.06.002
2009 Coleman,M, Gillanders,B, Connell,S, 2009, Dispersal and gene flow in the habitat-forming kelp, Ecklonia radiata: relative degrees of isolation across an east-west coastline, Marine and Freshwater Research, 60, 8, 802-809 10.1071/MF08268
2009 Gorman,D, Russell,B, Connell,S, 2009, Land-to-sea connectivity: Linking human-derived terrestrial subsidies to subtidal habitat change on open rocky coasts, Ecological Applications, 19, 5, 1114-1126 10.1890/08-0831.1
2008 Wernberg-Moller,T, Connell,S, 2008, Physical disturbance and subtidal habitat structure on open rocky coasts: Effects of wave exposure, extent and intensity, Journal of Sea Research, 59, 4, 237-248 10.1016/j.seares.2008.02.005
2008 Goodsell,P, Connell,S, 2008, Complexity in the relationship between matrix composition and inter-patch distance in fragmented habitats, Marine Biology, 154, 1, 117-125 10.1007/s00227-008-0906-2
2008 Valdivia,N, Stehbens,J, Hermelink,B, Connell,S, Molis,M, Wahl,M, 2008, Disturbance mediates the effects of nutrients on developing assemblages of epibiota, Austral Ecology, 33, 8, 951-962 10.1111/j.1442-9993.2008.01866.x
2008 Connell,S, Irving,A, 2008, Integrating ecology with biogeography using landscape characteristics: a case study of subtidal habitat across continental Australia, Journal of Biogeography, 35, 9, 1608-1621 10.1111/j.1365-2699.2008.01903.x
2008 Connell,S, Russell,B, Turner,D, Shepherd,A, Kildea,T, Miller,D, Airoldi,L, Cheshire,A, 2008, Recovering a lost baseline: missing kelp forests from a metropolitan coast, Marine Ecology-Progress Series, 360, 63-72 10.3354/meps07526
2008 Roberts,B, Connell,S, 2008, Detecting benthic responses to human-induced change: effectiveness of alternate taxonomic classification and indices, Marine Ecology-Progress Series, 358, 75-84 10.3354/meps07329
2007 Russell,B, Connell,S, 2007, Response of grazers to sudden nutrient pulses in oligotrophic versus eutrophic conditions, Marine Ecology-Progress Series, 349, 73-80 10.3354/meps07097
2007 Fowler-Walker,M, Connell,S, 2007, Habitat heterogeneity as a consequence of sub stratum-orientation and kelp-canopy: Relating interdependent responses to common patterns, Journal of Experimental Marine Biology and Ecology, 343, 1, 127-137 10.1016/j.jembe.2006.12.004
2007 Coleman,M, Vytopil,E, Goodsell,P, Gillanders,B, Connell,S, 2007, Diversity and depth-related patterns of mobile invertebrates associated with kelp forests, Marine and Freshwater Research, 58, 7, 589-595 10.1071/MF06216
2007 Glasby,T, Connell,S, Holloway,M, Hewitt,C, 2007, Nonindigenous biota on artificial structures: could habitat creation facilitate biological invasions?, Marine Biology, 151, 3, 887-895 10.1007/s00227-006-0552-5
2007 Johnston,E, Connell,S, Irving,A, Pile,A, Gillanders,B, 2007, Antarctic patterns of shallow subtidal habitat and inhabitants in Wilke' s Land, Polar Biology, 30, 6, 781-788 10.1007/s00300-006-0237-z
2006 Irving,A, Connell,S, 2006, Predicting understorey structure from the presence and composition of canopies: an assembly rule for marine algae, Oecologia, 148, 3, 491-502 10.1007/s00442-006-0389-0
2006 Fowler-Walker,M, Wernberg,T, Connell,S, 2006, Differences in kelp morphology between wave sheltered and exposed localities: morphologically plastic or fixed traits?, Marine Biology, 148, 4, 755-767 10.1007/s00227-005-0125-z
2006 Irving,A, Connell,S, 2006, Physical disturbance by kelp abrades erect algae from the understorey, Marine Ecology-Progress Series, 324, 127-137 10.3354/meps324127
2005 Fowler-Walker,M, Connell,S, Gillanders,B, 2005, Variation at local scales need not impede tests for broader scale patterns, Marine Biology, 147, 3, 823-831 10.1007/s00227-005-1605-x
2005 Goodsell,P, Connell,S, 2005, Disturbance initiates diversity in recruitment of canopy-forming algae: interactive effects of canopy-thinning and substratum availability, Phycologia, 44, 6, 632-639 10.2216/0031-8884(2005)44[632:DIDIRO]2.0.CO;2
2005 Russell,B, Gillanders,B, Connell,S, 2005, Proximity and size of neighbouring habitat affects invertebrate diversity, Marine Ecology-Progress Series, 296, IS, 31-38 10.3354/meps296031
2005 Drummond,S, Connell,S, 2005, Quantifying percentage cover of subtidal organisms on rocky coasts: a comparison of the costs and benefits of standard methods, Marine and Freshwater Research, 56, 6, 865-876 10.1071/MF04270
2005 Fowler-Walker,M, Gillanders,B, Connell,S, Irving,A, 2005, Patterns of association between canopy-morphology and understorey assemblages across temperate Australia, Estuarine Coastal and Shelf Science, 63, 1-2, 133-141 10.1016/j.ecss.2004.10.016
2005 Fowler-Walker,M, Connell,S, Gillanders,B, 2005, To what extent do geographic and associated environmental variables correlate with kelp morphology across temperate Australia?, Marine and Freshwater Research, 56, 6, 877-887 10.1071/MF05042
2005 Anderson,M, Connell,S, Gillanders,B, Diebel,C, Blom,W, Saunders,J, Landers,T, 2005, Relationships between taxonomic resolution and spatial scales of multivariate variation, Journal of Animal Ecology, 74, 4, 636-646 10.1111/j.1365-2656.2005.00959.x
2005 Connell,S, 2005, Assembly and maintenance of subtidal habitat heterogeneity: synergistic effects of light penetration and sedimentation, Marine Ecology-Progress Series, 289, 53-61 10.3354/meps289053
2005 Irving,A, Connell,S, Johnston,E, Pile,A, Gillanders,B, 2005, The response of encrusting coralline algae to canopy loss: an independent test of predictions on an Antarctic coast, Marine Biology, 147, 5, 1075-1083 10.1007/s00227-005-0007-4
2005 Russell,B, Connell,S, 2005, A novel interaction between nutrients and grazers alters relative dominance of marine habitats, Marine Ecology-Progress Series, 289, 5-11 10.3354/meps289005
2005 Russell,B, Elsdon,T, Gillanders,B, Connell,S, 2005, Nutrients increase epiphyte loads: broad-scale observations and an experimental assessment, Marine Biology, 147, 2, 551-558 10.1007/s00227-005-1571-3
2005 Goodsell,P, Connell,S, 2005, Historical configuration of habitat influences the effects of disturbance on mobile invertebrates, Marine Ecology-Progress Series, 299, 79-87 10.3354/meps299079
2005 Copertino,M, Connell,S, Cheshire,A, 2005, The prevalence and production of turf-forming algae on a temperate subtidal coast, Phycologia, 44, 3, 241-248 10.2216/0031-8884(2005)44[241:TPAPOT]2.0.CO;2
2004 Irving,A, Connell,S, Gillanders,B, 2004, Local complexity in patterns of canopy-benthos associations produces regional patterns across temperate Australasia, Marine Biology, 144, 2, 361-368 10.1007/s00227-003-1202-9
2004 Goodsell,P, Fowler-Walker,M, Gillanders,B, Connell,S, 2004, Variations in the configuration of algae in subtidal forests: Implications for invertebrate assemblages, Austral Ecology, 29, 3, 350-357 10.1111/j.1442-9993.2004.01372.x
2004 Irving,A, Connell,S, Elsdon,T, 2004, Effects of kelp canopies on bleaching and photosynthetic activity of encrusting coralline algae, Journal of Experimental Marine Biology and Ecology, 310, 1, 1-12 10.1016/j.jembe.2004.03.020
2004 Gorgula,S, Connell,S, 2004, Expansive covers of turf-forming algae on human-dominated coast: the relative effects of increasing nutrient and sediment loads, Marine Biology, 145, 3, 613-619 10.1007/s00227-004-1335-5
2003 Connell,S, 2003, Effects of habitat amount vs habitat configuration on persistance of small vs large populations inhabiting patchy landscapes, Records of the South Australian Museum Monograph Series, 7, 61-66
2003 Connell,S, 2003, Negative effects overpower the positive of kelp to exclude invertebrates from the understorey community, Oecologia, 137, 1, 97-103 10.1007/s00442-003-1312-6
2003 Irving,A, Fowler-Walker,M, Connell,S, 2003, Associations of forest-type with morphology of Ecklonia radiata and abundance of understorey algae, Transactions of the Royal Society of South Australia, 127, 2, 167-175
2003 Connell,S, 2003, The monopolization of understorey habitat by subtidal encrusting coralline algae: a test of the combined effects of canopy-mediated light and sedimentation, Marine Biology, 142, 6, 1065-1071 10.1007/s00227-003-1021-z
2002 Wellenreuther,M, Connell,S, 2002, Response of predators to prey abundance: separating the effects of prey density and patch size, Journal of Experimental Marine Biology and Ecology, 273, 1, 61-71 10.1016/S0022-0981(02)00145-4
2002 Irving,A, Connell,S, 2002, Sedimentation and light penetration interact to maintain heterogeneity of subtidal habitats: algal versus invertebrate dominated assemblages, Marine Ecology-Progress Series, 245, 83-91 10.3354/meps245083
2002 Fowler-Walker,M, Connell,S, 2002, Opposing states of subtidal habitat across temperate Australia: consistency and predictability in kelp canopy-benthic associations, Marine Ecology-Progress Series, 240, 49-56 10.3354/meps240049
2002 Irving,A, Connell,S, 2002, Interactive effects of sedimentation and microtopography on the abundance of subtidal turf-forming algae, Phycologia, 41, 5, 517-522 10.2216/i0031-8884-41-5-517.1
2002 Holloway,M, Connell,S, 2002, Why do floating structures create novel habitats for subtidal epibiota?, Marine Ecology-Progress Series, 235, 43-52 10.3354/meps235043
2002 Connell,S, 2002, Effects of a predator and prey on a foraging reef fish: implications for understanding density-dependent growth, Journal of Fish Biology, 60, 6, 1551-1561 10.1111/j.1095-8649.2002.tb02447.x
2002 Goodsell,P, Connell,S, 2002, Can habitat loss be treated independently of habitat configuration? Implications for rare and common taxa in fragmented landscapes, Marine Ecology-Progress Series, 239, 37-44 10.3354/meps239037
2001 Melville,A, Connell,S, 2001, Experimental effects of kelp canopies on subtidal coralline algae, Austral Ecology, 26, 1, 102-108 10.1111/j.1442-9993.2001.01089.pp.x
2001 Coleman,M, Connell,S, 2001, Weak effects of epibiota on the abundances of fishes associated with pier pilings in Sydney Harbour, Environmental Biology of Fishes, 61, 3, 231-239 10.1023/A:1011088631442
2001 Drabsch,S, Tanner,J, Connell,S, 2001, Limited infaunal response to experimental trawling in previously untrawled areas, ICES Journal of Marine Science, 58, 6, 1261-1271 10.1006/jmsc.2001.1105
2001 Glasby,T, Connell,S, 2001, Orientation and position of substrata have large effects on epibiotic assemblages, Marine Ecology-Progress Series, 214, 26 April, 127-135 10.3354/meps214127
2001 Saunders,R, Connell,S, 2001, Interactive effects of shade and surface orientation on the recruitment of spirorbid polychaetes, Austral Ecology, 26, 1, 109-115 10.1111/j.1442-9993.2001.01090.pp.x
2001 Connell,S, 2001, Predatory fish do not always affect the early development of epibiotic assemblages, Journal of Experimental Marine Biology and Ecology, 260, 1, 1-12 10.1016/S0022-0981(01)00246-5
2001 Connell,S, 2001, Urban structures as marine habitats: an experimental comparison of the composition and abundance of subtidal epibiota among pilings, pontoons and rocky reefs, Marine Environmental Research, 52, 2, 115-125 10.1016/S0141-1136(00)00266-X
2000 Connell,S, 2000, Is there satety-in-numbers for prey?, Oikos, 88, 3, 527-532 10.1034/j.1600-0706.2000.880308.x
2000 Connell,S, 2000, Encounter rates of a juvenile reef fish with small and predatory fishes, Copeia, 2000, 1, 36-41 10.1643/0045-8511(2000)2000[0036:EROAJR]2.0.CO;2
2000 Connell,S, 2000, Floating pontoons create novel habitats for subtidal epibiota, Journal of Experimental Marine Biology and Ecology, 247, 2, 183-194 10.1016/S0022-0981(00)00147-7
2000 Underwood,A, Chapman,M, Connell,S, 2000, Observations in ecology: You can't make progress on processes without understanding the patterns, Journal of Experimental Marine Biology and Ecology, 250, 1-2, 97-115 10.1016/S0022-0981(00)00181-7
1999 Connell,S, Lincoln-Smith,M, 1999, Depth and the structure of assemblages of demersal fish: experimental trawling along a temperate coast, Estuarine Coastal and Shelf Science, 48, 4, 483-495 10.1006/ecss.1998.0438
1999 Connell,S, Anderson,M, 1999, Predation by fish on assemblages of intertidal epibiota: effects of predator size and patch size, Journal of Experimental Marine Biology and Ecology, 241, 1, 15-29 10.1016/S0022-0981(99)00067-2
1999 Connell,S, Glasby,T, 1999, Do urban structures influence local abundance and diversity of subtidal epibiota? A case study from Sydney Harbour, Australia, Marine Environmental Research, 47, 4, 373-387 10.1016/S0141-1136(98)00126-3
1999 Connell,S, 1999, Effects of Surface Orientation on the Cover of Epibiota, Biofouling, 14, 3, 219-226 10.1080/08927019909378413
1999 Anderson,M, Connell,S, 1999, Predation by fish on intertidal oysters, Marine Ecology Progress Series, 187, 203-211 10.3354/meps187203
1999 Glasby,T, Connell,S, 1999, Urban structures as marine habitats, Ambio, 28, 7, 595-598
1998 Connell,S, Kingsford,J, 1998, Spatial, temporal and habitat-related variation in the abundance of large predatory fish at One Tree Reef, Australia, Coral Reefs, 17, 1, 49-57 10.1007/s003380050094
1998 Connell,S, 1998, Effects of predators on growth, mortality and abundance of a juvenile reef-fish: Evidence from manipulations of predator and prey abundance, Marine Ecology Progress Series, 169, 251-261 10.3354/meps169251
1998 Connell,S, Samoilys,M, Lincoln Smith,M, Leqata,J, 1998, Comparisons of abundance of coral-reef fish: Catch and effort surveys vs visual census, Austral Ecology, 23, 6, 579-586 10.1111/j.1442-9993.1998.tb00768.x
1998 Connell,S, 1998, Patterns of pisciviory by resident predatory reef fish at One Tree Reef, Great Barrier Reef, Marine and Freshwater Research, 49, 1, 25-30 10.1071/MF97034
1997 Connell,S, 1997, Exclusion of predatory fish on a coral reef: The anticipation, pre- emption and evaluation of some caging artefacts, Journal of Experimental Marine Biology and Ecology, 213, 2, 181-198 10.1016/S0022-0981(96)02773-6
1997 Connell,S, 1997, The relationship between large predatory fish and recruitment and mortality of juvenile coral reef-fish on artificial reefs, Journal of Experimental Marine Biology and Ecology, 209, 1-2, 261-278 10.1016/S0022-0981(96)02704-9
1996 Connell,S, 1996, Variations in mortality of a coral reef fish: Links with predator abundance, Marine Biology, 126, 2, 347-352 10.1007/BF00347459
1991 Connell,S, Jones,G, 1991, The influence of habitat complexity on postrecruitment processes in a temperate reef fish population, Journal of Experimental Marine Biology and Ecology, 151, 2, 271-294 10.1016/0022-0981(91)90129-K


Date Citation
2007 2007, Marine ecology, Oxford University Press, South Melbourne, 1-630

Book Chapters

Date Citation
2014 Wernberg,T, Russell,B, Thomsen,MS, Connell,SD, 2014, Marine biodiversity and climate change, Springer, pp. 181-187 10.1007/978-94-007-5784-4_80
2014 Russell,B, Connell,S, 2014, Ecosystem resilience and resistance to climate change, Springer, pp. 133-139 10.1007/978-94-007-5784-4_53
2014 Russell,BD, Connell,SD, 2014, Marine benthic productivity, Taylor and Francis, New York, pp. 767-771 10.1081/E-ENRW-120047577
2013 Falkenberg,L, Connell,S, Russell,B, 2013, Current and future eutrophication of nearshore marine environments: Causes, consequences and potential management strategies, pp. 215-238
2009 Harley,C, Connell,S, 2009, Shifts in abiotic variables and consequences for diversity, Springer, Heidelberg, Germany, pp. 257-268 10.1007/b76710_18
2009 Connell,S, Irving,A, 2009, The subtidal ecology of rocky coasts: Local-Regional-Biogeographic patterns and their experimental analysis, University of Chicago Press, USA, pp. 392-418
2009 Airoldi,L, Connell,S, Beck,M, 2009, The loss of natural habitats and the addition of artificial substrata, Springer, Heidelberg, Germany, pp. 269-280 10.1007/b76710_19
2007 Connell,S, Vanderklift,M, 2007, Negative interactions: The influence of predators and herbivores on prey and ecological systems, Oxford University Press, South Melbourne, pp. 72-100
2007 Connell,S, 2007, Water quality and the loss of coral reefs and kelp forests: alternative states and the influence of fishing, Oxford University Press, South Melbourne, pp. 556-568
2007 Connell,S, 2007, Subtidal temperate rocky habitats: habitat heterogenity at local to continental scales, Oxford University Press, South Melbourne, pp. 378-401
2005 Connell,S, Gillanders,B, 2005, Urban Ecological Communities 4: The marine and coastal environment, BioCity: Centre for Urban Habitats, Adelaide, pp. 298-318

Conference Papers

Date Citation
2016 Alleway,H, Thurstan,R, Lauer,P, Connell,S, 2016, Incorporating historical data into aquaculture planning, 3rd International Symposium on the Effects of Climate Change on the World's Oceans, PICES, Santos, BRAZIL 10.1093/icesjms/fsv191
2016 Bennett,S, Wernberg,T, Connell,S, Hobday,A, Johnson,C, Poloczanska,E, 2016, The 'Great Southern Reef': social, ecological and economic value of Australia's neglected kelp forests, 10th International Temperate Reefs Symposium, Perth, W.A. 10.1071/MF15232
2016 McSkimming,C, Russell,B, Tanner,J, Connell,S, 2016, A test of metabolic and consumptive responses to local and global perturbations: enhanced resources stimulate herbivores to counter expansion of weedy species, 10th International Temperate Reefs Symposium, Perth, W.A. 10.1071/MF14266


   Me writing a grant @ in the middle of the Great Australian Bight.

   My best ideas come whilst surrounded by the systems I study.




Grants with layperson titles:

  • ARC Discovery (2014-2016):  Ocean acidification propagates resource enrichment and its intensifying consumption in the web of life.
  • ARC Future Fellowship (2011-2013):  Global and local emissions boosts limiting resources for bottom-up global change.
  • ARC Linkage (2008 -2010): Managing global imprints with local activities to empower local managers.
  • ARC Linkage (2005-2007): Planning for coastal policy success by resolving land-to-sea linkages.
  • ARC Queen Elizabeth Fellowship II (2003-2005): Ecosystem decline in human-dominated landscapes.
  • ARC Discovery (2002-2005): Biogeography and ecology of southern kelp forests.
  • ARC Discovery (2000-2003): Experimental ecology of a subtidal rocky coast.






I teach the hard skills with love and passion.

1. Marine Biology.  I coordinate the BSc (Marine Biology).  We have taken students to China and East Timor on fully paid trips.

2. Statistics.  I coordinate the 3rd Year statistics course for biologists.  I receive unsolicited emails, such as "I just wanted to thank you for being a highly effective and thoughtful teacher during the first four weeks of RMEB III this semester.  I feel confident for the remainder of the course because of the way you presented the (relatively dry!) content and the way you listened to the students' needs"  (Student, 2017).

3. Writing.  Beautiful and nourishing reading comes from the effort we put into writing. Relative to maths, this one of the most under-valued skills in science.  Science writing is so dull and impenetrable that it sounds as if we are speaking in tongues.  Science communicators are employed to decode the babble.  I teach how to write clear and simple - the way to the truth - with a twist for a lively read.




I have graduated over 100 research students.  I work with students to design a project they can be passionate about, with an focus on the spirit of discovery and the needs of society.  My students have careers as marine biologists:

  1. Universities (national and international)
  2. Departments of conservation and wildlife
  3. Government water utilities
  4. Private consultancies
  5. Fisheries institutes
  6. Local councils
  7. Marine Parks






Date Role Membership Country
2013 Member The Environment Institute Australia

Committee Memberships

Date Role Committee Institution Country
2015 Member Habitat Enhancement Working Group Primary Industries South Australia Australia
2014 - 2016 Member International Temperate Reefs Symposium (Plymouth, UK) Plymouth University United Kingdom
2008 - 2016 Chair International Temperate Reefs Sympoium (Adelaide)
2002 - 2014 Advisory Board Member Scientific Working Group Department of Environment, Water and Natural Resources Australia
2001 - 2008 Board Member Wildlife Advsory Group Department of Environment and Hertiage Australia

Editorial Boards

Date Role Editorial Board Name Country
2010 - 2013 Associate Editor Aquatic Biology
2004 Associate Editor Marine Biology: international journal on life in oceans and coastal waters

Industry Partnerships

Date Engagement Type Partner Name
2013 Consultant SA Water


Date Institution Department Organisation Type Country
2006 - 2012 Department of Agriculture Great Australian Bight Fishery Australia
1994 - 1996 The Ecology Lab Australia
8313 4364
North Terrace
Darling Building
Room Number
1 17
Org Unit
Ecology and Environmental Science