Edward Green

Research Interests

Applied Mathematics Biological Mathematics Theoretical and Applied Mechanics

APrf Edward Green

Associate Professor

School of Mathematical Sciences

College of Science

I am an Associate Professor in Applied Mathematics in the School of Mathematical Sciences at the University of Adelaide, and a member of the Dynamics, Modelling and Computation (DMC) Group. My research interests are in mathematical biology and continuum mechanics (particularly fluid mechanics). Much of my work focuses on developing and analysing mathematical models to understand aspects of how tissues develop.

Current projects include:

  • Pattern formation in yeast colonies (with Dr Alex Tam, Associate Professor Ben Binder and Dr Daniel Netherwood (University of Adelaide), Professor Campbell Gourlay (University of Kent, UK) & Professor Vladimir Jiranek (University of Southampton, UK).
  • Modelling collective motion in animals, including locusts and sheep (with Dr Camille Buhl, Dr Stephan Leu and Dr Jack Maclean (University of Adelaide) and Professor Andrew Bernoff (Harvey Mudd College)

Date Position Institution name
2025 - ongoing Associate Professor University of Adelaide
2018 - 2024 Senior Lecturer in Applied Mathematics University of Adelaide
2011 - 2017 Lecturer in Applied Mathematics University of Adelaide
2010 - 2011 Research Assistant Professor University of Western Australia
2007 - 2009 Postdoctoral Fellow Ohio State University
2006 - 2006 Research Assistant University College London

Date Institution name Country Title
2006 University of Cambridge United Kingdom MA (Cantab)
2002 - 2006 University of Nottingham United Kingdom PhD
1999 - 2002 University of Cambridge United Kingdom BA (Hons)

Year Citation
2025 Georgiou, F., Buhl, C., Green, J. E. F., Lamichhane, B., & Thamwattana, N. (2025). Including population and environmental dynamic heterogeneities in continuum models of collective behaviour with applications to locust foraging and group structure. PLOS Computational Biology, 21(4), e1011469.
DOI WoS1 Europe PMC2
2025 Netherwood, D. J., Tam, A. K. Y., Gourlay, C. W., Knežević, T., Gardner, J. M., Jiranek, V., . . . Green, J. E. F. (2025). Accidental and Regulated Cell Death in Yeast Colony Biofilms. Bulletin of Mathematical Biology, 87(8, article no. 110), 110-1-110-23.
DOI
2025 Georgiou, F., Buhl, C., Green, J. E. F., Lamichhane, B., & Thamwattana, N. (2025). Including population and environmental dynamic heterogeneities in continuum models of collective behaviour with applications to locust foraging and group structure (vol 21, e1011469, 2025). PLOS COMPUTATIONAL BIOLOGY, 21(6), 1 page.
DOI
2024 Reoch, J. R., Stokes, Y. M., & Green, J. E. F. (2024). A two-phase thin-film model for cell-induced gel contraction incorporating osmotic effects. Journal of Mathematical Biology, 88(6), 61-1-61-42.
DOI
2024 Li, K., Green, E. J. F., Tronnolone, H., Tam, A. K. Y., Black, A. J., Gardner, J. M., . . . Binder, B. J. (2024). An off-lattice discrete model to characterise filamentous yeast colony morphology. PLoS Computational Biology, 20(11), e1012605-1-e1012605-20.
DOI Europe PMC3
2024 Brown, P. J., Green, J. E. F., Binder, B. J., & Osborne, J. M. (2024). Competing mechanisms for the buckling of an epithelial monolayer identified using multicellular simulation. Mathematical Biosciences, 380, 109367-1-109367-11.
DOI
2023 Li, K., Gallo, A. J., Binder, B. J., & Green, J. E. F. (2023). A mathematical model for nutrient–limited uniaxial growth of a compressible tissue. Journal of Theoretical Biology, 575, 111631-1-111631-12.
DOI Scopus1 WoS1 Europe PMC1
2023 Hopwood, M. J., Harding, B., Green, J. E. F., & Dyson, R. J. (2023). Thin film extensional flow of a transversely isotropic viscous fluid. Physical Review Fluids, 8(11), 113301-1-113301-33.
DOI
2022 Reoch, J. R., Stokes, Y. M., & Green, J. E. F. (2022). A mathematical model for cell-induced gel contraction incorporating osmotic effects. Journal of Mathematical Biology, 84(5), 31-1-31-42.
DOI Scopus2 WoS2 Europe PMC2
2022 Georgiou, F., Buhl, C., Green, J. E. F., Lamichhane, B., & Thamwattana, N. (2022). Modelling foraging competition between solitarious and gregarious organisms in increasingly heterogeneous environments. Journal of Insect Physiology, 143, 104443.
DOI Scopus6 WoS6 Europe PMC5
2022 Reoch, J. R., Stokes, Y. M., & Green, J. E. F. (2022). A mathematical model for cell-induced gel contraction incorporating osmotic effects. Journal of Mathematical Biology, 84, 31.
DOI
2022 Tam, A. K. Y., Harding, B., Green, J. E. F., Balasuriya, S., & Binder, B. J. (2022). Thin-film lubrication model for biofilm expansion under strong adhesion. Physical Review E, 105(1), 1-15.
DOI Scopus6 WoS6 Europe PMC6
2022 Brown, P. J., Green, J. E. F., Binder, B. J., & Osborne, J. M. (2022). Publisher Correction: A rigid body framework for multicellular modeling. Nature Computational Science, 2(1), 59.
DOI
2021 Georgiou, F., Buhl, J., Green, J. E. F., Lamichhane, B., & Thamwattana, N. (2021). Erratum: Modelling locust foraging: How and why food affects group formation (PLoS Comput Biol (2021) 17: 7 (e1008353) DOI: 10.1371/journal.pcbi.1008353). PLoS Computational Biology, 17(12), e1009695.
DOI Scopus1 WoS1
2021 Brown, P., Green, E., Binder, B., & Osborne, J. (2021). A rigid body framework for multi-cellular modelling.
DOI Europe PMC1
2021 Gallo, A. J., Tronnolone, H., Green, J. E., & Binder, B. J. (2021). Modelling uniaxial non-uniform yeast colony growth: comparing an agent-based model and continuum approximations. Journal of Theoretical Biology, 523, 110715-1-110715-13.
DOI Scopus1 WoS1 Europe PMC1
2021 Georgiou, F., Buhl, C., Green, J. E. F., Lamichhane, B., & Thamwattana, N. (2021). Modelling locust foraging: How and why food affects group formation. PLOS Computational Biology, 17(7), e1008353.
DOI Scopus16 WoS15 Europe PMC9
2021 Brown, P. J., Green, J. E. F., Binder, B. J., & Osborne, J. M. (2021). A rigid body framework for multicellular modeling. Nature Computational Science, 1(11), 754-766.
DOI Scopus10 WoS11 Europe PMC8
2020 Georgiou, F., Buhl, J., Green, J. E. F., Lamichhane, B., & Thamwattana, N. (2020). Modelling locust foraging: How and why food affects hopper band formation.
DOI
2019 Arnold, D. J., Stokes, Y. M., & Green, J. E. F. (2019). Particle-laden thin-film flow in helical channels with arbitrary shallow cross-sectional shape. Physics of Fluids, 31(7), 073305-1-073305-17.
DOI Scopus5 WoS5
2019 Tam, A., Green, J. E. F., Balasuriya, S., Tek, E. L., Gardner, J. M., Sundstrom, J. F., . . . Binder, B. J. (2019). A thin-film extensional flow model for biofilm expansion by sliding motility. Proceedings of the Royal Society A: Mathematical, Physical and Engineering Sciences, 475(2229), 20190175-1-20190175-21.
DOI Scopus11 WoS12 Europe PMC10
2018 Tam, A., Green, J., Balasuriya, S., Tek, E., Gardner, J., Sundstrom, J., . . . Binder, B. (2018). Nutrient-limited growth with non-linear cell diffusion as a mechanism for floral pattern formation in yeast biofilms. Journal of Theoretical Biology, 448, 122-141.
DOI Scopus16 WoS17 Europe PMC12
2018 Tronnolone, H., Tam, A., Szenczi, Z., Green, J., Balasuriya, S., Tek, E., . . . Binder, B. (2018). Diffusion-limited growth of microbial colonies. Scientific Reports, 8(5992), 5992-1-5992-11.
DOI Scopus32 WoS28 Europe PMC23
2018 Dyson, R. J., & Green, J. E. F. (2018). Editorial: Mechanics in biology. Anziam Journal, 59(4), 413-415.
DOI
2018 Holloway, C., Cupples, G., Smith, D., Green, J., Clarke, R., & Dyson, R. (2018). Influences of transversely isotropic rheology and translational diffusion on the stability of active suspensions. Royal Society Open Science, 5(8), 1-18.
DOI Scopus7 WoS6 Europe PMC1
2018 Green, J., Whiteley, J., Oliver, J., Byrne, H., & Waters, S. (2018). Pattern formation in multiphase models of chemotactic cell aggregation. Mathematical Medicine and Biology: A Journal of the IMA, 35(3), 319-346.
DOI Scopus7 WoS7 Europe PMC4
2018 Dini, S., Binder, B., & Green, J. (2018). Understanding interactions between populations: individual based modelling and quantification using pair correlation functions. Journal of Theoretical Biology, 439, 50-64.
DOI Scopus15 WoS13 Europe PMC14
2017 Arnold, D., Stokes, Y., & Green, J. (2017). Thin-film flow in helically wound shallow channels of arbitrary cross-sectional shape. Physics of Fluids, 29(1), 013102-1-013102-11.
DOI Scopus8 WoS7
2016 Dyson, R., Green, J., Whiteley, J., & Byrne, H. (2016). An investigation of the influence of extracellular matrix anisotropy and cell-matrix interactions on tissue architecture. Journal of Mathematical Biology, 72(7), 1775-1809.
DOI Scopus25 WoS25 Europe PMC12
2016 Psaltis, S., Green, J., Farrell, T., Lawson, B., & Simpson, J. (2016). Optimising Cheese Brining Times. ANZIAM Journal, 56, 57-94.
DOI WoS1
2016 Cui, X., Dini, S., Dai, S., Bi, J., Binder, B., Green, J., & Zhang, H. (2016). A mechanistic study on tumour spheroids formation in thermosensitive hydrogels: experiments and mathematical modelling. RSC Advances, 6(77), 73282-73291.
DOI Scopus29 WoS22
2016 Snelling, E., Seymour, R., Green, J., Meyer, L., Fuller, A., Haw, A., . . . Maloney, S. (2016). A structure-function analysis of the left ventricle. Journal of Applied Physiology, 121(4), 900-909.
DOI Scopus10 WoS9 Europe PMC9
2016 Dini, S., Binder, B., Fischer, S., Mattheyer, C., Schmitz, A., Stelzer, E., . . . Green, J. (2016). Identifying the necrotic zone boundary in tumour spheroids with pair-correlation functions. Journal of the Royal Society Interface, 13(123), 20160649-1-20160649-11.
DOI Scopus31 WoS29 Europe PMC22
2015 Arnold, D., Stokes, Y., & Green, J. (2015). Thin-film flow in helically-wound rectangular channels of arbitrary torsion and curvature. Journal of Fluid Mechanics, 764, 76-94.
DOI Scopus11 WoS10
2014 Agnew, D., Green, J., Brown, T., Simpson, M., & Binder, B. (2014). Distinguishing between mechanisms of cell aggregation using pair-correlation functions. Journal of Theoretical Biology, 352, 16-23.
DOI Scopus47 WoS43 Europe PMC37
2014 Davies, K., Green, J., Bean, N., Binder, B., & Ross, J. (2014). On the derivation of approximations to cellular automata models and the assumption of independence. Mathematical Biosciences, 253(1), 63-71.
DOI Scopus9 WoS8 Europe PMC7
2013 Green, J., Bassom, A., & Friedman, A. (2013). A mathematical model for cell-induced gel compaction in vitro. Mathematical Models & Methods in Applied Sciences, 23(1), 127-163.
DOI Scopus4 WoS4
2010 Green, J., Waters, S., Whiteley, J., Edelstein-Keshet, L., Shakesheff, K., & Byrne, H. (2010). Non-local models for the formation of hepatocyte-stellate cell aggregates. Journal of Theoretical Biology, 267(1), 106-120.
DOI Scopus37 WoS39 Europe PMC28
2009 Matzavinos, A., Kao, C., Green, J., Sutradhar, A., Miller, M., & Friedman, A. (2009). Modeling oxygen transport in surgical tissue transfer. Proceedings of the National Academy of Sciences of the United States of America, 106(29), 12091-12096.
DOI Scopus16 WoS14 Europe PMC15
2009 Green, J., Smith, F., & Ovenden, N. (2009). Flow in a multi-branching channel with compliant walls. Journal of Engineering Mathematics, 64(4), 353-365.
DOI Scopus6 WoS6
2009 Green, J., Waters, S., Shakesheff, K., & Byrne, H. (2009). A mathematical model of liver cell aggregation in vitro. Bulletin of Mathematical Biology, 71(4), 906-930.
DOI Scopus26 WoS21 Europe PMC12
2008 Green, J., & Friedman, A. (2008). The extensional flow of a thin sheet of incompressible, transversely isotropic fluid. European Journal of Applied Mathematics, 19(3), 225-257.
DOI Scopus16 WoS13
- Psaltis, S., Green, J. E. F., Farrell, T., Lawson, B., & Simpson, J. L. (2016). Optimising Cheese Brining Times. ANZIAM Journal, 56, 57.
DOI
- Georgiou, F. H., Lamichhane, B., Thamwattana, N., Buhl, J., & Green, E. (2022). A numerical scheme for non-local aggregation with non-linear diffusion and approximations of social potential. ANZIAM Journal, 62, C242-C255.
DOI
- Broadbridge, T. P., Green, J. E. F., Preston, S. P., Fadai, N. T., & Maclean, J. (2025). Food purchase data reveals the locations of London’s ‘food deserts’. PLOS Complex Systems, 2(11), e0000072.
DOI

Year Citation
2009 Ma, X., Green, J. E. F., Gooch, K. J., Jansen, R. M., & Hart, R. T. (2009). Finite element modeling of entangled collagen fiber gels with randomly oriented fibers. In Proceedings of the ASME Summer Bioengineering Conference 2009, SBC2009 (pp. 1041-1042). USA: AMER SOC MECHANICAL ENGINEERS.
DOI
2003 Green, J. E. F., Byrne, H. M., Waters, S. L., & Shakesheff, K. M. (2003). Mathematical modelling in tissue engineering. In European Cells and Materials Vol. 6 (pp. 38).

Year Citation
- Tam, A., Balasuriya, S., Green, E., & Binder, B. (n.d.). Understanding floral pattern formation in yeast biofilms. Poster session presented at the meeting of Unknown Conference.
DOI

Year Citation
2024 Li, K., Green, E., Tronnolone, H., Tam, A., Black, A., Gardner, J., . . . Binder, B. (2024). An off-lattice discrete model to characterise filamentous yeast colony morphology.
DOI

2013-2016 Australian Research Council Discovery Early Career Researcher Award

2023-2026 Australian Research Council Discovery Project (DP230100406)

2023- Analytics for the Australian Grains Industry

Date Role Research Topic Program Degree Type Student Load Student Name
2025 Principal Supervisor Mathematical modelling of plaque calcification in atherosclerosis Master of Philosophy Master Full Time Miss Faith Caren Thando Penn Sawers
2025 Principal Supervisor Mathematical modelling of plaque calcification in atherosclerosis Master of Philosophy Master Full Time Miss Faith Caren Thando Penn Sawers
2024 Co-Supervisor Mathematical modelling of the evolution of ecologically scaffolded life cycles Master of Philosophy Master Full Time Mr Daniel John Fraterman
2024 Co-Supervisor Mathematical modelling of the evolution of ecologically scaffolded life cycles Master of Philosophy Master Full Time Mr Daniel John Fraterman
2023 Co-Supervisor Mathematical modelling and spatial quantification of yeast colony growth Doctor of Philosophy Doctorate Full Time Mr Kai Li
2023 Co-Supervisor Mathematical modelling and spatial quantification of yeast colony growth Doctor of Philosophy Doctorate Full Time Mr Kai Li
2021 Principal Supervisor Modelling and Quantification of Food Deserts Doctor of Philosophy under a Jointly-awarded Degree Agreement with Doctorate Part Time Miss Tayla Paige Broadbridge
2021 Principal Supervisor Modelling and Quantification of Food Deserts Doctor of Philosophy Doctorate Full Time Miss Tayla Paige Broadbridge

Date Role Research Topic Program Degree Type Student Load Student Name
2018 - 2020 Co-Supervisor Modelling One-Dimensional Non-Uniform Growth with Applications to Cylindrical Yeast Colonies Master of Philosophy Master Full Time Mr Anthony John Gallo
2017 - 2021 Principal Supervisor Transversely isotropic thin film flows Doctor of Philosophy under a Jointly-awarded Degree Agreement with Doctorate Full Time Mr Matthew Hopwood
2017 - 2024 Principal Supervisor A New Approach to Agent Based Multi-Cellular Modelling Doctor of Philosophy Doctorate Part Time Mr Phillip James Brown
2016 - 2019 Co-Supervisor Mathematical Modelling of Pattern Formation in Yeast Biofilms Doctor of Philosophy Doctorate Full Time Mr Alexander Tam
2014 - 2018 Co-Supervisor Spatial Quantification and Mathematical Modelling of Tissue Development Doctor of Philosophy Doctorate Full Time Mr Saber Dini
2013 - 2016 Co-Supervisor Thin-Film Flow in Helical Channels Doctor of Philosophy Doctorate Full Time Mr David John Arnold
2013 - 2020 Co-Supervisor Mathematical Modelling of The Complex Mechanics of Biological Gels Doctor of Philosophy Doctorate Part Time Mr James Richard Reoch

Date Role Committee Institution Country
2022 - ongoing Member Equity, Diversity and Inclusion Committee Australian Mathematical Society Australia

Date Role Editorial Board Name Institution Country
2023 - ongoing Associate Editor PLoS Complex Systems University of Adelaide Australia
2017 - ongoing Associate Editor Australia and New Zealand Industrial and Applied Mathematics (ANZIAM) Journal University of Adelaide Australia

Date Office Name Institution Country
2013 - 2016 Chair SA Branch of ANZIAM Australia

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